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EET 95 [ECU]    Theobroma cacao

Derivation: Estacion Experimental Tropical.
Location: E.E.T. Pichilingue, ECU.

Notes:  Some EET clones are imported genotypes which have been given the duplicate identifier [IMP], [POU] for kown Pound collections and [MIG] for known SAN MIGUEL clones.
 

Synonyms: TENGUEL 33 [ECU], ETS 95 [ECU]

 NOTES

Some EET clones are imported genotypes which have been given the duplicate identifier [IMP], [POU] for kown Pound collections and [MIG] for known SAN MIGUEL clones.

 RECOMMENDED PLANTING MATERIAL    Disclaimer

 INIAP, 2020
Recommended for planting in:
Ecuador
Recommended by: INIAP
Year: 1970's (Current)
 Reference    View Associated Data

 PEDIGREE DETAILS

 Parents:
NACIONAL  x VEN. AMARILLO TYPE   Ecuador - Pichilingue, 1987
NACIONAL  x Unknown   Lockwood, G., 1973
 Mother to:
EET 451 [ECU]    Ecuador - Pichilingue, 1987
EET 460 [ECU]    Ecuador - Pichilingue, 1987

 GENOME DATA LINKS

 Cornejo, O.E. et al., 2018  Reference
BioProject: PRJNA486011   NCBI   ENA FASTQ 
Project: PRJNA486011   EVA (project)   EVA (analysis) 
 
Run: SRR7774211   NCBI   ENA FASTQ 
Experiment: SRX4629613
BioSample: SAMN09934440
SampleSource: Ecuador

 HELD IN

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Current List   ICQC,R Website
• Accession: RUQ 414     ICQC,R 
   - Local Name: EET 95 [ECU]
   - Year of Accession: 1993
List received in 2012   Reference
• Local Name: EET-95
   - Year of Accession: 1996
List received in 1990   Reference
• Local Name: EET 95 [ECU]
List received in 2018   Reference
• Accession: 10273
   - Local Name: EET-95
   - Year of Accession: 1965
List received in 1987   Reference
• Local Name: EET 95 [ECU]
   - Year of Accession: 1974
List received in 2011   Reference
• Local Name: EET-95
List received in 2012   Reference
• Local Name: EET - 95
List received in 2012   Reference
• Local Name: EET95
List received in 1994   Reference
• Local Name: EET 95 [ECU]
List received in 1994   Reference
• Local Name: EET 95 [ECU]
List received in 1995   Reference
• Local Name: EET 95 [ECU]
   - Year of Accession: 1987
List received in 2009   Reference
• Local Name: EET - 95
List received in 2011   Reference
• Local Name: EET 95
   - Year of Accession: 1966

 DISEASE

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 Ceratocystis
Tolerant
 INIAP, 2020
Intermediate
 Morera, J. & Mora, A., 1990
Intermediate
 Enriquez, G.A. & Soria, V.J., 1967
 Frosty Pod Rot
Moderately Susceptible
 INIAP, 2020
Moderately Susceptible, 93.00 % Infection
Note: External Severety = 3.0 and Internal Severety = 3.4
 Phillips, W., 1996
Moderately Susceptible, 93.00 % Infection
Note: External Severety = 3.0 and Internal Severety = 3.4
 Phillips, W., 1997
 Phytophthora
Moderate Resistance, sp. P. palmivora (Pod).
Note: Unripe, Attached and not wounded
 Morera, J., 1993
Moderately Susceptible, sp. P. palmivora (Pod).
 Phillips, W., 1997
Resistant, 18.00 % Infection, sp. P. palmivora (Pod).
Note: Unripe and not wounded
 Lopez-Baez, O. et al., 1996
 Witches Broom
Susceptible
 INIAP, 2020
Resistant
 Morera, J. & Mora, A., 1990
Susceptible
 Enriquez, G.A. & Soria, V.J., 1967

 FLAVOUR

 INIAP, 2020
Flavour: medium fruit/floral aroma, highly adaptable
 Reference

 YIELD

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 Frances Bekele & Gillian Bidaisee, 2016
Pod Index: 27.2 (pods per kg dried beans)
 Reference
 Costa Rica - CATIE, 1981
Seed Index: 77.5 (dried beans per 100g)
Productivity: high
 Reference
 Trinidad - ICGT, 1998
Pod Index: 26.7 (pods per kg dried beans)
 Reference
 INIAP, 2020
Yield: 1750 kg ha-1 yr-1
 Reference

 BEAN

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 Bartley, B.G.D., 1970
Size: large
 Reference
 Frances Bekele & Gillian Bidaisee, 2016
Colour: dark purple
Shape: elliptical
Number: 46.0
Total Wet Weight: 88.1 g
Cotyledon Dry Weight: 0.80 g
Cotyledon Length: 24.5 mm
Cotyledon Width: 13.0 mm
 Reference
 Costa Rica - CATIE, 1981
Colour Dark Purple: Present
Colour Light Purple: Absent
Colour Grey: Absent
Colour White: Absent
Colour Mottled: Absent
Number: 36.3
Maximum Number: 50
Cotyledon Wet Weight: 1.93 g
Cotyledon Dry Weight: 1.29 g
Cotyledon Length: 25.1 mm
Cotyledon Width: 12.3 mm
Cotyledon Thickness: 9.0 mm
 Reference
 Enriquez, G.A. & Soria, V.J., 1967
Colour Purple: 100 %
Shape: flat/round
Length: 26.8 mm
Width: 14.2 mm
Thickness: 9.4 mm
Cotyledon Wet Weight: 2.29 g
Cotyledon Dry Weight: 1.54 g
 Reference
 Trinidad - ICGT, 1998
Colour: dark purple
Shape: elliptical
Number: 45.0
Total Wet Weight: 84.3 g
Total Dry Weight: 52.5 g
Cotyledon Dry Weight: 0.83 g
Cotyledon Length: 24.6 mm
Cotyledon Width: 13.0 mm
Cotyledon Thickness: 8.3 mm
 Reference
 INIAP, 2020
Dry Weight: 1.30 g
 Reference
 Morera, J. & Mora, A., 1990
Colour Dark Purple: 100 %
Cotyledon Wet Weight: 2.29 g
Cotyledon Dry Weight: 1.54 g
Cotyledon Length: 26.8 mm
Cotyledon Width: 14.2 mm
Cotyledon Thickness: 9.4 mm
 Reference

 BUTTERFAT

 Trinidad - CRU, 1998
Butterfat Content: 51.60 %
Testa: 6.10 %
 Reference
 INIAP, 2020
Butterfat Content: 50.00 %
 Reference

 FRUIT

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 Bartley, B.G.D., 1970
Size: medium
 Reference
 Frances Bekele & Gillian Bidaisee, 2016
Shape: elliptical
Basal Constriction: moderate
Apex Form: obtuse
Rugosity: moderate
Length: 176.0 mm
Breadth: 90.0 mm
Colour: anthocyanin absent
Note: Pod wall hardness > 2.0 MPa
 Reference
 Costa Rica - CATIE, 1981
Basal Constriction: very slight
Apex Form: slightly acute
Rugosity: moderate
Husk Thickness: 11.15 mm (13.9 - 8.4)
Length: 189.0 mm
Breadth: 92.0 mm
Colour: absent anthocyanin
Weight: 690.0 g
Pods Per Tree: 7
Hardness: moderate
Length Breadth Ratio: 2.05
Note: ratio of distances between ridges within a pair and between ridges of two different pairs = 0.66
 Reference
 Enriquez, G.A. & Soria, V.J., 1967
Shape: cundeamor
Husk Thickness: 15.9 mm
Length: 179.0 mm
Breadth: 93.0 mm
Colour: yellow
Hardness: very hard
Length Breadth Ratio: 1.79
 Reference
 Trinidad - ICGT, 1998
Shape: angoleta
Basal Constriction: moderate
Apex Form: obtuse
Rugosity: moderate
Husk Thickness: 16.40 mm
Size: large
Length: 168.5 mm
Breadth: 88.4 mm
Colour: anthocyanin absent
Husk Weight: 372.70 g
Placenta Weight: 6.16 g
Hardness: hard
Length Breadth Ratio: 1.90
 Reference
 Morera, J. & Mora, A., 1990
Shape: cundeamor
Rugosity: intense
Husk Thickness: 15.9 mm
Length: 179.0 mm
Breadth: 93.0 mm
Colour: yellow
Hardness: hard
Length Breadth Ratio: 1.92
 Reference
 Morera, J. et al., 1991
Shape: amelonado
Colour: green
 Reference

 PHENOTYPING

 Fiona Lahive, 2017
Accession: RUQ 414
Light Saturated Photosynthetic Rate: 3.74 μmol m2 s-1
Stomatal Conductance: 0.07 mol m2 s-1
Transpiration Rate: 0.92 mmol m2 s-1
Intrinsic Water Use Efficiency: 62.03 μmol mol-1
Instantaneous Water Use Efficiency: 4.52 mmol mol-1
Flush Size: 5
Leaf Area: 329.80 cm2
Leaf Dry Weight: 2.01 g
Specific Leaf Area: 165.10 cm2 g-1
 Reference

 FLUSH

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 Costa Rica - CATIE, 1981
Colour: reddish brown
 Reference
 Enriquez, G.A. & Soria, V.J., 1967
Colour: slight anthocyanin
 Reference
 Trinidad - ICGT, 1992
Colour: intermediate anthocyanin
 Reference
 Morera, J. & Mora, A., 1990
Colour: green
 Reference

 LEAF

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 Enriquez, G.A. & Soria, V.J., 1967
Length to Width Ratio: 2.37
Basal Angle: 143.0 °
Apical Angle: 96.0 °
 Reference
 Trinidad - ICGT, 1992
Distance from Base to Widest Part: 128 mm
Length: 258.0 mm
Width: 100.0 mm
Length to Width Ratio: 2.60
Basal Angle: 112.0 °
Apical Angle: 78.1 °
Petiole Length: 10 mm
Pulvinus Length: 17 mm
 Reference
 Morera, J. & Mora, A., 1990
Length to Width Ratio: 2.37
Basal Angle: 143.0 °
Apical Angle: 96.0 °
 Reference

 FLOWER

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 Frances Bekele & Gillian Bidaisee, 2016
Ligule Colour: moderate anthocyanin
Stamen Colour: slight anthocyanin
Sepal Length: 8.38 mm
Ligule Width: 2.76 mm
Ovule Number: 46
Style Length: 2.69 mm
Pedicel Colour: red
 Reference
 Costa Rica - CATIE, 1981
Ligule Colour: intense anthocyanin (apical)
Stamen Colour: intense anthocyanin
Sepal Colour: greyish green
Sepal Width: 2.28 mm
Sepal Length: 9.08 mm
Ligule Length: 7.85 mm
Ligule Width: 2.53 mm
Ovary Colour: anthocyanin absent (apical)
Ovary Width: 0.98 mm
Ovary Length: 1.32 mm
Style Length: 2.62 mm
Staminode Length: 6.94 mm
Pedicel Colour: pastel red
Note: Translation of Methuen colour codes.
 Reference
 Enriquez, G.A. & Soria, V.J., 1967
Bud Colour: slight anthocyanin
Ligule Colour: moderate anthocyanin
Stamen Colour: intense anthocyanin
Guide Line Length: 3.10 mm
Sepal Colour: anthocyanin absent
Sepal Width: 2.60 mm
Sepal Length: 8.70 mm
Sepal Length to Width Ratio: 3.28
Ligule Length: 3.70 mm
Ligule Width: 3.00 mm
Ligule Length to Width Ratio: 1.22
Ribbon Length: 3.90 mm
Ribbon Colour: anthocyanin absent
Ovary Colour: anthocyanin absent (apical), anthocyanin absent (basal)
Ovary Width: 1.40 mm
Ovary Length: 1.90 mm
Ovary Length to Width Ratio: 1.35
Ovule Number: 51
Style Colour: anthocyanin absent (apical), anthocyanin absent (basal)
Style Length: 2.90 mm
Staminode Colour: intense anthocyanin
Staminode Length: 7.50 mm
Pedicel Length: 20.10 mm
Pedicel Colour: intense anthocyanin, intense anthocyanin (ab), moderate anthocyanin (basal)
Hairs on Upper Surface of Sepal: present
Pigmented Glands on Lower Surface of Sepal: present
Horns at Base of Guide Lines: absent
Pigmented Spot on Pollen Sac: absent
 Reference
 Trinidad - ICGT, 1998
Ligule Colour: slight anthocyanin
Stamen Colour: intense anthocyanin
Sepal Length: 8.43 mm
Ligule Width: 2.76 mm
Ovule Number: 37
Style Length: 2.77 mm
Pedicel Colour: very slight anthocyanin
 Reference
 Morera, J. & Mora, A., 1990
Bud Colour: green
Ligule Colour: green - white
Stamen Colour: moderate anthocyanin
Guide Line Length: 3.10 mm
Sepal Colour: white
Sepal Width: 2.60 mm
Sepal Length: 8.70 mm
Ligule Length: 3.70 mm
Ligule Width: 3.00 mm
Ribbon Length: 3.90 mm
Ribbon Colour: white
Ovary Width: 1.40 mm
Ovary Length: 1.90 mm
Ovule Number: 51
Style Colour: white (apical), white (basal)
Style Length: 2.90 mm
Staminode Colour: yellow
Staminode Length: 7.50 mm
Pedicel Length: 20.10 mm
Pedicel Colour: yellow, yellow (ab), green - white (basal)
Hairs on Upper Surface of Sepal: present
Pigmented Glands on Lower Surface of Sepal: present
Horns at Base of Guide Lines: absent
 Reference
 Morera, J. et al., 1991
Self Compatibility: self compatible
 Reference

INIAP (2020) Venta de semillas y plantas [Sale of seeds and plants]. INIAP Website
Ecuador - Pichilingue (1987) Nomina de clones y numero de plantas existentes en la coleccion internacional de germoplasma de cacao, EET, Pichilingue.. Unpublished report, E.E.T. Pichilingue, Ecuador. pp. 18.
Lockwood, G. (1973) Report on a visit to cocoa centres in the USA, the Caribbean and Latin America for acquisition of germplasm with particular reference to cocoa swollen shoot disease.. Unpublished report, British Research Team, Cocoa Research Institite, Ghana. pp. 23.
Cornejo, O.E., Muh-Ching Yee, Victor Dominguez, Mary Andrews, Alexandra Sockell, Erika Strandberg, Donald Livingstone III, Conrad Stack, Alberto Romero, Pathmanathan Umaharan, Stefan Royaert, Nilesh R. Tawari, Pauline Ng, Osman Gutierrez, Wilbert Phillips, Keithanne Mockaitis, Carlos D. Bustamante & Juan C. Motamayor (2018) Population genomic analyses of the chocolate tree, Theobroma cacao L., provide insights into its domestication process. Commun Biol 1, 167
Mars Center for Cocoa Science (2012) Accession list for the Mars Center for Cocoa Science.. Jean-Philippe Marelli, personal communication, February 2012.
Colombia - ICA Palmira (1990) List of clones held at ICA Palmira.. F. Ocampo Rojas, ICA, Colombia.
Allan Mata (2018) Accession list for the International Cacao Collection at CATIE (IC3).. Personal communication.
Dominican Republic - CDC (1987) Informe de los trabajos de indentificacion clonal realizados por el Dr. B.G.Bartley durante su visita a la Estacion "Mata Larga".. Unpublished of the Comision del Cacau, Dominican Republic. pp 14
IDIAF (2011) Cocoa accessions held in the Dominican Institute for Agriculture and Forestry Research (IDIAF).. M. Ventura, personal communication, September 2011.
EET - Pichilingue (2012) Accessions planted in EET - Pichilingue, Ecuador.. Freddy Amores and Ignacio Sotomayor C, personal communication, February 2012.
CRIG (2012) Cocoa accessions held in the Cocoa Research Institute of Ghana (CRIG).. Francis Kwame Padi, personal communication, February 2012.
Guatemala - Los Brilliantes (1994) Lista de clones existentes en la Estacion de Investigacion y Produccion Agricola "Los Brillantes" a Agosto 1994.. M.T. Reyes Barrios, EIAP Guatemala. Personal Communication.
Honduras - CEDEC (1994) List of accessions in CEDEC, La Masica, Honduras.. J.A. Sanchez, FHIA, Honduras. Personal Communication.
Lopez-Baez, O., Cueto-Moreno, J. & Fraire-Vazquez, G. (1995) Progressos en el mejoramiento genetico del cacao Theobroma cacao L. en Mexico.. In: Proceedings of the International Workshop on Cocoa Breeding Strategies, Kuala Lumpur, 1994. INGENIC, Reading, U.K. pp. 18-25.
Luis F. García Carrión (2009) Catalogo de Cultivares de Cacao del Peru.
CRU (2011) ICG,T accessions.. CRU website.
Morera, J. & Mora, A. (1990) Banco de Germoplasma de Cacao del CATIE.. Information supplied on computer diskette by CATIE, Turrialba, Costa Rica.
Enriquez, G.A. & Soria, V.J. (1967) Cacao Cultivars Register.. IICA Teaching and Research Centre, Costa Rica
Phillips, W. (1996) Cocoa cultivar reaction when innoculated with Moniliophthora roreri. . Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication
Phillips, W. (1997) Results of the artificial inoculation of Moniliophthora rorei using the Aspersion method.. Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication
Morera, J. (1993) Reaction of clones to Phytophthora palmivora. . J. Morera, CATIE, Costa Rica. Personal Communication.
Phillips, W. (1997) Results of the artificial inoculation of Phytophthora palmivora using the Paper Disc method.. Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication
Lopez-Baez, O., Fraire, V.G. & Cueto, M.J. (1996) Mejoramiento genetico para la resistencia a Phytophthora en Mexico.. Proceedings of the International Workshop on the Contribution of Disease Resistance to Cocoa Variety Improvement, INGENIC, Salvador, 24th-26th September, 1996. pp.51-55.
Frances Bekele & Gillian Bidaisee (2016) Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago.. Unpublished data on fruit and flower morphology supplied as an Excel spreadsheet by Frances Bekele, April 2016.
Costa Rica - CATIE (1981) Genetic Resources of Cacao. A Catalogue of the CATIE Collection.. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.
Trinidad - ICGT (1998) Flower and fruit descriptions.. Unpublished data supplied on computer diskette by International Cocoa Genebank, Trinidad, University of the West Indies
Bartley, B.G.D. (1970) A preliminary catalogue of the cacao germplasm held at the University of the West Indies, Trinidad.. University of the West Indies, St. Augustine, Trinidad.
Trinidad - CRU (1998) Fat Analysis Data.. Data supplied on computer diskette by F. Bekele, CRU, Trinidad.
Morera, J., Paredes, A. & Mora, A. (1991) Germoplasma de cacao en el CATIE entre 1947 y 1991, Programa II: Generacion y transferencia de tecnologia.. IICA, San Jose, Costa Rica, pp. 49.
Fiona Lahive (2017) Phenotypic data from climate change trials.. Personal communication.
Trinidad - ICGT (1992) Flower, fruit and leaf descriptors.. Unpublished data supplied on computer diskette by International Cocoa Genebank, Trinidad, University of the West Indies, Trinidad.