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TSA 656    Theobroma cacao

 Shripat, C. (1996)
The Cocoa Breeding Programme of the Ministry of Agriculture, Land and Marine Resources, Trinidad. Presented at the International Workshop on The Utilisation of Genetic Resources of the ICGT, June, 1996. Cocoa Research Unit, Trinidad.

Derivation: Trinidad Selected Amazons.
Collected by Cocoa Board.
Location: Trinidad and Tobago.

Notes:  Selections from populations of Amazons.

Shripat, 1996a [SHR96A]. Each Trinidad clone was labelled with a code composed of certain letters as a prefix (TS, TSA, TSH, TSAN, TSHN) followed by a number which runs consecutively from 500 upwards regardless of the letters. Thus, there are no two new clones with the same number.

(see also Bartley, 1997c [BAR97C] ).


 Lopes, Uilson Vanderlei et al., 2011
Recommended clone in:
Brazil (Bahia)
Year: 1998-2014
 Yamada, Milton Macoto et al., 2010
Recommended clone in:
Brazil (Bahia)
Year: 1990


SCA 6  x IMC 67    Lockwood, G. & Gyamfi, M.M.O., 1979
 Father to:
CEPEC 25    Brazil - CEPEC, 1987


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List received in 2012   Reference
• Accession: 282
   - Local Name: TSA 656
   - Year of Accession: 1980
List received in 1987   Reference
• Local Name: TSA 656
   - Year of Accession: 1979
List received in 2015   Reference
• Accession: TSA 656 Quadra 8
   - Local Name: TSA 656
List received in 2012   Reference
• Local Name: TSA-656
List received in 2018   Reference
• Accession: 22090
   - Local Name: TSA-656
   - Year of Accession: 2007


Susceptible (Branch).
Note: Based on mycelium growth and the production of perithecia..
 Stela Dalva Vieira Midlej Silva, 2003
 Witches Broom
 Pires, J.L. et al., 1996


 Brazil - CEPEC, 1987
Rugosity: intense


 Fiona Lahive, 2017
Accession: RUQ 1016
Light Saturated Photosynthetic Rate: 5.13 μmol m2 s-1
Stomatal Conductance: 0.11 mol m2 s-1
Transpiration Rate: 1.16 mmol m2 s-1
Intrinsic Water Use Efficiency: 54.92 μmol mol-1
Instantaneous Water Use Efficiency: 4.45 mmol mol-1
Flush Size: 5
Leaf Area: 199.20 cm2
Leaf Dry Weight: 1.34 g
Specific Leaf Area: 150.70 cm2 g-1


 Frances Bekele & Gillian Bidaisee, 2022
Colour: moderate anthocyanin


 Brazil - CEPEC, 1987
Self Compatibility: self incompatible

Shripat, C. (1996) The Cocoa Breeding Programme of the Ministry of Agriculture, Land and Marine Resources, Trinidad. Presented at the International Workshop on The Utilisation of Genetic Resources of the ICGT, June, 1996. Cocoa Research Unit, Trinidad.
Lopes, Uilson Vanderlei, Wilson Reis Monteiro, José Luis Pires, Didier Clement, Milton Macoto Yamada & Karina Peres Gramacho (2011) Cacao Breeding in Bahia, Brazil - Strategies and Results. Crop Breeding and Applied Biotechnology 11: 73–81
Yamada, Milton Macoto, Fábio Gelape Faleiro, Brena Faria Santos & Mônica Menezes Passinho (2010) RELAÇÃO ENTRE INCOMPATIBILIDADE E VARIABILIDADE GENÉTICA COM BASE EM MARCADORES RAPD EM CLONES DE CACAUEIRO [Relationship between Incompatibility and Genetic Variability Based on RAPD Markers in Cacao Clones]. Agrotrópica (Itabuna) 22 (1): 11–16
Lockwood, G. & Gyamfi, M.M.O. (1979) The CRIG cocoa germplasm collection with notes on codes used in the breeding programme at Tafo and elsewhere. Technical Bulletin, No. 10, Cocoa Research Institute, Tafo, Ghana. pp. 62.
Brazil - CEPEC (1987) List of clones in the CEPEC, Bahia, Brazil collection. Computer printout from CEPEC. Supplied by B.G.D. Bartley, CEPEC, Brazil.
CEPEC (2012) Accession list for the CEPEC (Bahia, Brazil) collection. Dr. José Luis Pires, personal communication, February 2012.
Almeida, C.M.V.C. de, Barriga, J.P., Machado, P.F.R. & Bartley, B.G.D. (1987) Evolucao do programa de conservacao dos recursos geneticos de cacau na Amazonia Brasileira. CEPLAC/DEPEA, Boletim Tecnico, Belem, Brasil 5: pp.108.
Almeida, Caio Márcio Vasconcellos Cordeiro de, Francisco Antônio Neto, Acácio de Paula, Silva, José Raymundo Pires Medeiros & Antonio de Almeida Lima (2015) RECURSOS GENÉTICOS DE CACAUEIRO EM RONDÔNIA: RETROSPECTIVA HISTÓRICA, ORIGEM E INVENTÁRIO [Genetic Resources of Cacao in Rondônia: Historical Overview, Origin and Inventory]. Agrotrópica (Itabuna) 27 (2): 93–124
Mars Center for Cocoa Science (2012) Accession list for the Mars Center for Cocoa Science. Jean-Philippe Marelli, personal communication, February 2012.
Allan Mata (2018) Accession list for the International Cacao Collection at CATIE (IC3). Personal communication.
Stela Dalva Vieira Midlej Silva (2003) Ensaios Para Avaliação do Cacaueiro à Murcha-de-Ceratocystis na Bahia, Brasil. In: Proceedings of the 14th International Cocoa Research Conference, Accra, Ghana (October 2003). Cocoa Producers' Alliance, Lagos, Nigeria.
Pires, J.L., Monteiro, W.R., Pinto, L.R.M. & Luz, E.D.M.N. (1996) Resistance to Witches Broom - Evaluation of genotypes from different origins. In: Proceedings of the 12th International Cocoa Research Conference, Salvador, Brasil 1996 Cocoa Producers' Alliance (in press).
Fiona Lahive (2017) Phenotypic data from climate change trials. Personal communication.
Frances Bekele & Gillian Bidaisee (2022) Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.