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IMC 23 Group Name Theobroma cacao

Pound, F.J. (1938)
Cacao and Witchbroom Disease of South America with notes on other species of Theobroma. Archives of Cocoa Research 1: pp. 26-71.

Derivation: Iquitos Mixed Calabacillo (Mid Channel).
Collected by Pound.
Location: Iquitos, Peru.

Pound, 1938a [POU38A]. Main type - very large lagarta calabacillo pods, oval, very smooth, normal blanco. Seeds large, flat, losenge shaped. 'Hybrid group'-large pods, somewhat warty at stem end but smooth and round like calabacillo at the apex. Third group-small, oval, warty pods, dense flowering. Bartley, 1995a [BAR95A]. The common features of this group of clones are the high ovule numbers (65+) and a specific type of anthocyanin pigmentation in the stamens and base of the corolla. Posnette, 1945a [POS45A]. Petioles are of a type intermediate between those of Nanay (see NA) and Parinari (see PA) types; the constriction is usually short and narrow.

Notes: Pound, 1938a [POU38A]. Pods probably collected from at least two groups of trees which were free of Witches' Broom.

Bartley, 1995a [BAR95A]. They are probably the progenies of two trees, later propagated as the clones P 18 [POU] and P 21 [POU].

Pound, 1943b [POU43B]. 112 plants were planted at Marper farm.

Bartley, 1997a, [BAR97A] The progeny of the individual clones is not documented and it is important not to consider the IMC clones as one homogeneous group.

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Synonyms: IQUITOS MIXED 23

PEDIGREE DETAILS

Putative Half-Sibs:: Group P27 Zhang, D. et al., 2009

Reference

Zhang, D., Boccara, M., Motilal, L., Mischke S., Johnson, E.S., Butler, D.R., Bailey, B. & Meinhardt, L. 2009
Molecular characterization of an earliest cacao (Theobroma cacao L.) collection from Upper Amazon using microsatellite DNA markers. Tree Genetics & Genomes. doi:10.1007/s11295-009-0212-2; IMC 10; IMC 105; IMC 107; IMC 11; IMC 12; IMC 13; IMC 16; IMC 18; IMC 27; IMC 30; IMC 33; IMC 38; IMC 42; IMC 44; IMC 47; IMC 53; IMC 55; IMC 58; IMC 59; IMC 6; IMC 60; IMC 61; IMC 63; IMC 65; IMC 68; IMC 71; IMC 76; IMC 77; IMC 78; IMC 85; IMC 9; IMC 96; IMC 98

POPULATION

Motamayor, J.C. et al., 2008: 'Iquitos' Population; Subgroup: 'IMC I' Reference

Reference

Motamayor, J.C., Lachenaud, P., da Silva e Mota, J.W., Loor, R., Kuhn, D.N., Brown, J.S. & Schnell, R.J. 2008
Geographic and Genetic Population Differentiation of the Amazonian Chocolate Tree (Theobroma cacao L). PLoS ONE 3(10): e3311. doi:10.1371/journal.pone.0003311
Population SSR

HELD IN

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Brazil, Comissão Executiva do Plano da Lavoura Cacaueira/Centro de Pesquisas do Cacau (CEPLAC/CEPEC): List received in 2012 Reference

Reference

CEPEC 2012
Accession list for the CEPEC (Bahia, Brazil) collection. Dr. José Luis Pires, personal communication, February 2012.; • Accession: 559; - Local Name: IMC 23; - Year of Accession: 1985
Cote d'Ivoire, Centre National de Recherche Agronomique (CNRA): List received in 1989 Reference

Reference

Cote d'Ivoire - IRCC 1989
Liste des clones en collection. Unpublished Report, IRCC Bingerville, Cote d'Ivoire, Doc. No. 192. D. Clement, IRCC, Cote d'Ivoire. Personal Communication.
Notes

List of clones grouped as West African Amelonados, Lower Amazons, Venezuelan collections, French Guiana collections, Ecuadorian collections, local Trinitarios, other Trinitarios, upper Amazons and Criollos. Notes on origin, date and place of origin and nature of the material (cuttings, budwood, seed).; • Local Name: IMC 23; - Year of Accession: 1977
Ecuador, Estation Experimental Tropical Pichilingue: List received in 2012 Reference

Reference

EET - Pichilingue 2012
Accessions planted in EET - Pichilingue, Ecuador. Freddy Amores and Ignacio Sotomayor C, personal communication, February 2012.; • Local Name: IMC - 23
Malaysia, Agricultural Research Centre (ARC): List received in 1990 Reference

Reference

Malaysia - ARC Tuaran 1990
List of clones held in the Dept. of Agriculture's collection, at Sabah, Tuaran, Malaysia. E.B. Tay, Malaysian Cocoa Board, Malaysia. Personal Communication.; • Local Name: IMC 23
Togo, Institut de Recherches du Cafe et du Cacao (IRCC): List received in 1991 Reference

Reference

Togo - IRCC Togo 1991
Collection des clones de cacaoyers presents au Togo. Unpublished report, IRCC Togo, p.1.; • Local Name: IMC 23
Trinidad and Tobago, International Cocoa Genebank, Trinidad (ICG,T): List received in 2011 Reference

Reference

CRU 2011
ICG,T accessions. CRU website.
Notes

Website visited 12th April 2011

Link Unavailable [http://sta.uwi.edu/cru/icgt-accessions.asp]; • Local Name: IMC 23

DISEASE

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Phytophthora: Susceptible, sp. P. palmivora (Pod).; Note: Unripe, Unattached and not wounded; Iwaro, D., 1998

Reference

Iwaro, D. 1998
Evalation of accessions for Black Pod. Personal Communication 30th October 1998. Data sent by email from Dr. Iwaro.
Notes

Trinidad - CRU, University of the West Indies Datafile: DIS_PHDS - Disease reaction to Phytophthora Experimental conditions: Inoculation techinique used - spray method. Inoculum concentration: 100,000 zoospores/ml. Incubated for four days. Character Data: The following descriptions are used to segregate clones according to reaction: no visible symptoms = highly resistant to penetration, 1-5 localised lesions = resistant, 6-15 localised lesions = moderately resistant, >15 localised lesions = partially resistant/resistant to spread of lesion alone, 1-5 expanding lesions = partially resistant/resistant to penetration alone, 6-15 expanding lesions = moderately susceptible, >15 expanding lesions = susceptible, coalesced lesions = highly susceptible Disease - Phytophthora palmivora, P. megakarya (Local Names: Black Pod, Podredumbre negra, Pourriture Brune, Podridao parda).

Disease
Cocoa Swollen Shoot Virus: Tolerant; Lockwood, G., 1986

Reference

Lockwood, G. 1986
Report on cocoa breeding & seed production. Unpublished report for the Government of Ghana, Cocoa Research Institute of Ghana, CDC (UK.). pp. 23.
Notes

Ghana - CRIG Datafile: DIS_SSDS - Disease reaction to Swollen shoot Descriptors: Notes on cocoa germplasm, breeding programmes, clones that have been lost from Q6 Environment: No details given Germplasm: No details given

Disease
Vascular Streak Dieback: 42.60 % Infection (Field Reaction).; Note: 14.3% of trees with <30% infected branches, 19.3% of trees with 30 to 70% infected branches, 9.0% of trees with >70% infected branches.. Sime Darby Plantations, Devon Estate.; Lam, K.S., 1992

Reference

Lam, K.S. 1992
Performance of imported clones in Sime Darby Plantations. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Notes

Malaysia - Sime Darby Plantations, Selangor Agronomic characteristics - Sime Darby Plantations, Disease - Oncobasidium theobromae (local names: Vascular Streak Dieback) Datafile: DIS_VSDS - Disease reaction to VSD SIMDARDS MYS - Sime Darby Plantations Germplasm: Descriptor information is presented for 129 of the clones held at the Plantations' Devon and Merlimau Estates. Environment: No details. Experimental Conditions - Sime Darby: Trial BK/GP/1 and BK/GP/2 Location: Devon Estate Date planted: April 1986 Pod count: Feb 1988 to Jan 1991 Pod analysis: Nov 1988 to Jan 1991 Planting distance: 9' x 10' No. of replicates: 3 No. of trees/plot: 16 Expt. Design: Randomised Complete Block Design Trial BK/GP/3 Location: Merlimau Estate Date planted: March 1988 Planting distance: 9' x 10' No. of replicates: 2 No. of trees/plot: 16 Expt. Design: Randomised Complete Block Design

Disease Yield Bean; 43.33 % Infection; Note: 18 plants sampled..; Subali, S., 1992

Reference

Subali, S. 1992
MARDI's results on performance of imported primary cocoa clones. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Notes

Malaysia - MARDI Datafile: MARDI_DS MYS - MARDI Germplasm: Agronomic and morphological characteristics of some of the clones held by MARDI. Environment: No details.

Disease

YIELD

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Malaysia - Applied Agriculture, 1992: Seed Index: 101.0 (dried beans per 100g); Pod Index: 21.7 (pods per kg dried beans); Reference

Reference

Malaysia - Applied Agriculture 1992
Data from Applied Agriculture Research Unit (AARU). In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October 1992, Kuala Lumpur, Malaysia: pp. 4.
Notes

Malaysia - Applied Agriculture Research Datafile: AARU__DS MYS - Applied Agriculture Germplasm: Agronomic characteristics of 120 of the clones held at Applied Agriculture Research. The clones were field budded on 1982 seedlings planting between 1984 and 1987. Environment: No details given.

Yield Bean
Malaysia - BAL Plantations, 1991: Seed Index: 103.0 (dried beans per 100g); Pod Index: 20.8 (pods per kg dried beans); Reference

Reference

Malaysia - BAL Plantations 1991
Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.
Notes

Malaysia - BAL Plantations Datafile: BAL2__DS MYS - BAL Plantations (update) Environment: Trials are planted on Table Estate, Tawau. Trial CA81 - planted August 1987 in 3x6 tree plots, 3 replicates randomised blocks, trees spaced at 3.8m x 2.4 m (1,096 trees/ha). Trial CA88 - similar design, planted August 1988. Soil type is fairly fertile young volcanic basalt. Average rainfall is c. 2100 mm/year well distributed, the driest month being February with about 150 mm. Descriptors: Cocoa butter analysis:- samples of a minimum of 2 kg were fermented in net bags placed in the fermentation heap and subsequently fast (i.e. continuously) dried. Determinations were carried out by MacRobertson Foods Pte Ltd. (a division of Cadbury-Schweppes Ltd.) in Singapore using a Nuclear Magnetic Resonance machine.

Yield Bean Butterfat Flower
Frances Bekele & Gillian Bidaisee, 2022: Pod Index: 26.1 (pods per kg dried beans); Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Lam, K.S., 1992: Seed Index: 117.3 (dried beans per 100g); Pod Index: 31.6 (pods per kg dried beans); Reference

Reference

Lam, K.S. 1992
Performance of imported clones in Sime Darby Plantations. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Notes

Malaysia - Sime Darby Plantations, Selangor Agronomic characteristics - Sime Darby Plantations, Disease - Oncobasidium theobromae (local names: Vascular Streak Dieback) Datafile: DIS_VSDS - Disease reaction to VSD SIMDARDS MYS - Sime Darby Plantations Germplasm: Descriptor information is presented for 129 of the clones held at the Plantations' Devon and Merlimau Estates. Environment: No details. Experimental Conditions - Sime Darby: Trial BK/GP/1 and BK/GP/2 Location: Devon Estate Date planted: April 1986 Pod count: Feb 1988 to Jan 1991 Pod analysis: Nov 1988 to Jan 1991 Planting distance: 9' x 10' No. of replicates: 3 No. of trees/plot: 16 Expt. Design: Randomised Complete Block Design Trial BK/GP/3 Location: Merlimau Estate Date planted: March 1988 Planting distance: 9' x 10' No. of replicates: 2 No. of trees/plot: 16 Expt. Design: Randomised Complete Block Design

Disease Yield Bean

BEAN

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Malaysia - Applied Agriculture, 1992: Number: 46.3; Dry Weight: 0.99 g; Reference

Reference

Malaysia - Applied Agriculture 1992
Data from Applied Agriculture Research Unit (AARU). In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October 1992, Kuala Lumpur, Malaysia: pp. 4.
Notes

Malaysia - Applied Agriculture Research Datafile: AARU__DS MYS - Applied Agriculture Germplasm: Agronomic characteristics of 120 of the clones held at Applied Agriculture Research. The clones were field budded on 1982 seedlings planting between 1984 and 1987. Environment: No details given.

Yield Bean
Malaysia - BAL Plantations, 1991: Number: 49.4; Total Dry Weight: 48.1 g; Dry Weight: 0.97 g; Note: 58 plants sampled; Reference

Reference

Malaysia - BAL Plantations 1991
Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.
Notes

Malaysia - BAL Plantations Datafile: BAL2__DS MYS - BAL Plantations (update) Environment: Trials are planted on Table Estate, Tawau. Trial CA81 - planted August 1987 in 3x6 tree plots, 3 replicates randomised blocks, trees spaced at 3.8m x 2.4 m (1,096 trees/ha). Trial CA88 - similar design, planted August 1988. Soil type is fairly fertile young volcanic basalt. Average rainfall is c. 2100 mm/year well distributed, the driest month being February with about 150 mm. Descriptors: Cocoa butter analysis:- samples of a minimum of 2 kg were fermented in net bags placed in the fermentation heap and subsequently fast (i.e. continuously) dried. Determinations were carried out by MacRobertson Foods Pte Ltd. (a division of Cadbury-Schweppes Ltd.) in Singapore using a Nuclear Magnetic Resonance machine.

Yield Bean Butterfat Flower
Bartley, B.G.D., 1970: Size: medium; Reference

Reference

Bartley, B.G.D. 1970
A preliminary catalogue of the cacao germplasm held at the University of the West Indies, Trinidad. University of the West Indies, St. Augustine, Trinidad.
Notes

Trinidad - University of the West Indies Datafile: ICGT2_DS TTO - UWI Cacao Collection Germplasm: Passport descriptors for approximately 1000 accessions of mainly South American and Trinidadian origin. Environment: No details. Descriptors: Six descriptors for fruit, seed and compatability characteristics. Character Data: Combinations used FXC = forastero x criollo, TX = trinitario cross

Bean Fruit Flower
Frances Bekele & Gillian Bidaisee, 2022: Colour: purple; Shape: ovate; Number: 51.0; Total Wet Weight: 53.8 g; Cotyledon Dry Weight: 0.75 g; Cotyledon Length: 21.8 mm; Cotyledon Width: 11.8 mm; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Lam, K.S., 1992: Number: 41.9; Dry Weight: 0.85 g; Reference

Reference

Lam, K.S. 1992
Performance of imported clones in Sime Darby Plantations. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Notes

Malaysia - Sime Darby Plantations, Selangor Agronomic characteristics - Sime Darby Plantations, Disease - Oncobasidium theobromae (local names: Vascular Streak Dieback) Datafile: DIS_VSDS - Disease reaction to VSD SIMDARDS MYS - Sime Darby Plantations Germplasm: Descriptor information is presented for 129 of the clones held at the Plantations' Devon and Merlimau Estates. Environment: No details. Experimental Conditions - Sime Darby: Trial BK/GP/1 and BK/GP/2 Location: Devon Estate Date planted: April 1986 Pod count: Feb 1988 to Jan 1991 Pod analysis: Nov 1988 to Jan 1991 Planting distance: 9' x 10' No. of replicates: 3 No. of trees/plot: 16 Expt. Design: Randomised Complete Block Design Trial BK/GP/3 Location: Merlimau Estate Date planted: March 1988 Planting distance: 9' x 10' No. of replicates: 2 No. of trees/plot: 16 Expt. Design: Randomised Complete Block Design

Disease Yield Bean
Palaniappan, S. & Shuhaimi Shamsuddin, 1992: Number: 45.4; Dry Weight: 1.26 g; Reference

Reference

Palaniappan, S. & Shuhaimi Shamsuddin 1992
Early evaluation of some cocoa germplasm at Pusut Perkhidmatan Pertanian Tun Razak. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Notes

Malaysia - FELDA, Pusat Perkhidmatan Pertanian Tun Razak, Peninsula Malaysia Datafile: FELDA_DC MYS - Felda PPPTR Germplasm: Agronomic and morphological characteristics of some of the clones held at Pusat Perkhidmatan Pertanian Tun Razak. The clones were obtained from the Malaysian Cocoa Growers Council Joint Importation Scheme, MARDI and the Department of Agriculture, Peninsular Malaysia. Environment: The clones are planted in single plots of 18 plants/clone (6x3) at Pusat Perkhidmatan Pertanian Tun Razak, Sg. Tekam in inland Peninsular Malaysia. The clones were planted as they were obtained and therefore are of varying age. Character data: Seed dry weight: corrected to 6% moisture after oven drying at 105'C for 24 hours.

Bean Fruit

BUTTERFAT

Malaysia - BAL Plantations, 1991: Butterfat Content: 56.60 %; Maximum Butterfat Content: 59.35 %; Minimum Butterfat Content: 55.08 %; Note: 13 plants sampled; Reference

Reference

Malaysia - BAL Plantations 1991
Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.
Notes

Malaysia - BAL Plantations Datafile: BAL2__DS MYS - BAL Plantations (update) Environment: Trials are planted on Table Estate, Tawau. Trial CA81 - planted August 1987 in 3x6 tree plots, 3 replicates randomised blocks, trees spaced at 3.8m x 2.4 m (1,096 trees/ha). Trial CA88 - similar design, planted August 1988. Soil type is fairly fertile young volcanic basalt. Average rainfall is c. 2100 mm/year well distributed, the driest month being February with about 150 mm. Descriptors: Cocoa butter analysis:- samples of a minimum of 2 kg were fermented in net bags placed in the fermentation heap and subsequently fast (i.e. continuously) dried. Determinations were carried out by MacRobertson Foods Pte Ltd. (a division of Cadbury-Schweppes Ltd.) in Singapore using a Nuclear Magnetic Resonance machine.

Yield Bean Butterfat Flower
Frances Bekele & Gillian Bidaisee, 2022: Testa: 6.20 %; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower

FRUIT

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Bartley, B.G.D., 1970: Size: large; Reference

Reference

Bartley, B.G.D. 1970
A preliminary catalogue of the cacao germplasm held at the University of the West Indies, Trinidad. University of the West Indies, St. Augustine, Trinidad.
Notes

Trinidad - University of the West Indies Datafile: ICGT2_DS TTO - UWI Cacao Collection Germplasm: Passport descriptors for approximately 1000 accessions of mainly South American and Trinidadian origin. Environment: No details. Descriptors: Six descriptors for fruit, seed and compatability characteristics. Character Data: Combinations used FXC = forastero x criollo, TX = trinitario cross

Bean Fruit Flower
Frances Bekele & Gillian Bidaisee, 2022: Shape: obovate; Basal Constriction: slight; Apex Form: obtuse; Rugosity: moderate; Length: 147.0 mm; Breadth: 80.0 mm; Colour: anthocyanin absent; Hardness: moderate; Note: Hardness > 1.6 and <= 2.0 Mpa; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Brazil - CEPEC, 1987: Rugosity: intense; Reference

Reference

Brazil - CEPEC 1987
List of clones in the CEPEC, Bahia, Brazil collection. Computer printout from CEPEC. Supplied by B.G.D. Bartley, CEPEC, Brazil.
Notes

Brazil - CEPEC, Itabuna, Bahia Datafile: CEPEC_DS BRA - CEPEC 1 PROGENY_ Germplasm: 564 accessions mainly of Brazilian origin. Environment: No details. Descriptors: Information on compatability pod characteristics and type of cocoa is given.

Bean Fruit Flower
Palaniappan, S. & Shuhaimi Shamsuddin, 1992: Length: 179.6 mm; Breadth: 91.5 mm; Weight: 649.4 g; Length Breadth Ratio: 2.00; Reference

Reference

Palaniappan, S. & Shuhaimi Shamsuddin 1992
Early evaluation of some cocoa germplasm at Pusut Perkhidmatan Pertanian Tun Razak. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Notes

Malaysia - FELDA, Pusat Perkhidmatan Pertanian Tun Razak, Peninsula Malaysia Datafile: FELDA_DC MYS - Felda PPPTR Germplasm: Agronomic and morphological characteristics of some of the clones held at Pusat Perkhidmatan Pertanian Tun Razak. The clones were obtained from the Malaysian Cocoa Growers Council Joint Importation Scheme, MARDI and the Department of Agriculture, Peninsular Malaysia. Environment: The clones are planted in single plots of 18 plants/clone (6x3) at Pusat Perkhidmatan Pertanian Tun Razak, Sg. Tekam in inland Peninsular Malaysia. The clones were planted as they were obtained and therefore are of varying age. Character data: Seed dry weight: corrected to 6% moisture after oven drying at 105'C for 24 hours.

Bean Fruit

FLUSH

Allen, J.B., 1988: Colour: slight anthocyanin; Reference

Reference

Allen, J.B. 1988
Geographical variation and population biology in wild Theobroma cacao. PhD. Thesis, University of Edinburgh, U.K. pp. 197.
Notes

Colombia, Ecuador - Amazon region Datafile: LCT___DC ECU - LCTAP Fruit and seed characteristics LCT2__DS LCTAP Habit and flower characteristics LCT___DS ECU - LCTAP Field and genebank data Germplasm: Details of 234 LCT-EEN and EBC accessions from material collected in the Amazon region of Ecuador and R. Caqueta, Colombia and several of Pound's collections held at the San Carlos genebank. Environment: Accessions were either described at the time of collection or after planting in the field genebank at the San Carlos site of the INIAP Napo-Payamino Experimental Station in Napo Province. In the field genebank, a maximum of 10 seedlings for each collection were planted in a single row and budwood collections were planted as a row of up to 5 plants. Trees were spaced at 3m within the row and 4m between rows. Meteorological data are given in the table as mean daily maximum and mean daily minimum temperature and mean total rainfall per month. Soil analyses were carried out in the Department of Forestry and Natural Resources, Edinburgh University, UK. The table shows the means of duplicate analyses of five samples taken from the genebank site and the forest surrounding the genebank. Soil data: pH 4.2 Org. Mat. (%) 9.4 N (Total, %) 0.3 P (Total, µg/g) 702 Ca (meq/100g) 5.19 Mg (meq/100g) 1.60 K (meq/100g) 0.48 Ca/Mg (ratio) 3.24 Ca+Mg/K (ratio) 14.1 Meteorological data: Temperature (°C) Rainfall (mm) Daily Max. Daily Min. January 30.9 21.2 116.2 February 29.1 20.6 272.3 March 29.8 21.2 260.0 April 29.9 21.3 327.5 May 29.9 21.0 287.9 June 27.9 20.4 308.1 July 28.1 20.0 254.1 August 29.3 20.3 262.5 September 30.3 20.5 282.4 October 30.6 21.1 257.1 November 30.8 21.1 361.2 December 31.0 21.4 154.8 Descriptors: The coding of the descriptors, and the methodology of recording, is based on the IBPGR standardised format, Engels et al., 1980 [ENG80A] and Engels, 1981 [ENG81A]. However, pod and bean data were recorded from a sample of 1-5 pods per collection, rather than the recommended sample of 10-20 pods due to limited availability of pods.

Flush Flower
Frances Bekele & Gillian Bidaisee, 2022: Colour: slight anthocyanin; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower

LEAF

Trinidad - ICGT, 1992: Distance from Base to Widest Part: 113 mm; Length: 244.0 mm; Width: 87.0 mm; Length to Width Ratio: 2.80; Basal Angle: 132.5 °; Apical Angle: 74.3 °; Petiole Length: 7 mm; Pulvinus Length: 16 mm; Reference

Reference

Trinidad - ICGT 1992
Leaf descriptors. Unpublished data supplied on computer diskette by International Cocoa Genebank, Trinidad, University of the West Indies, Trinidad.
Notes

Trinidad - International Cocoa Genebank, CRU, University of the West Indies.
Flower descriptor data is presented for 620 clones, fruit descriptor data of 333 and seed characters of 118 clones are partly described. Leaf descriptor data is presented for 229 clones. The germplasm is mainly of Peruvian, Ecuadorian and Colombian origin. Environment: Each clone is represented in the collection by up to 16 trees planted 1.8 m apart in a block design. Shading is provided by Erythrina sp. (spaced 60m apart) and bananas (spaced 4m apart). Fertilizer, N-P-K (15:10:5), is provided twice a year at 250g per tree. Cyclic irrigation is performed from February to May (10 l of water per plant/week). Soil type group -33 A 1 Soil Series - Cunupia fine sandy clay with restricted internal drainage Lithology - silty clay alluvium Drainage - imperfect Slope category - 0°-2° Meterological data are given in this table as mean maximum, mean minimum and overall mean temperature and mean total rainfall per month over the period 1961-1990. Month Temp. (°C) Rainfall Max Min Mean (mm) January 30.1 20.8 24.6 71.2 February 30.2 20.8 25.0 43.3 March 31.2 21.4 25.7 36.6 April 31.8 22.3 26.5 51.1 May 31.6 23.3 26.9 116.5 June 30.6 23.3 26.4 252.1 July 30.8 23.0 26.2 266.3 August 31.2 22.9 26.4 250.1 September 31.5 22.8 26.5 202.5 October 31.4 22.82 6.41 99.3 November 30.8 22.4 25.9 227.8 December 30.2 21.5 25.2 155.6 Descriptors: The character selection is based on the IBPGR list (IBPGR, 1981 [IBP81A]). Where descriptor states present real measurements, e.g. pod length in cm, the data are given as the mean of all the measurements ± the standard deviation. For some descriptors, such as Leaf, flush colour, the phenotypic expression of the character is coded on a scale from 0 to 9. On this scale '0' represents the lowest or weakest expression of a characteristic and '9' represents the highest or strongest. Leaf and Flower Characteristics p.300-307

Leaf

FLOWER

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Allen, J.B., 1988: Stamen Colour: slight anthocyanin; Sepal Colour: anthocyanin absent; Sepal Length: 8.00 mm; Ovary Colour: anthocyanin absent (apical); Pedicel Colour: anthocyanin absent; Reference

Reference

Allen, J.B. 1988
Geographical variation and population biology in wild Theobroma cacao. PhD. Thesis, University of Edinburgh, U.K. pp. 197.
Notes

Colombia, Ecuador - Amazon region Datafile: LCT___DC ECU - LCTAP Fruit and seed characteristics LCT2__DS LCTAP Habit and flower characteristics LCT___DS ECU - LCTAP Field and genebank data Germplasm: Details of 234 LCT-EEN and EBC accessions from material collected in the Amazon region of Ecuador and R. Caqueta, Colombia and several of Pound's collections held at the San Carlos genebank. Environment: Accessions were either described at the time of collection or after planting in the field genebank at the San Carlos site of the INIAP Napo-Payamino Experimental Station in Napo Province. In the field genebank, a maximum of 10 seedlings for each collection were planted in a single row and budwood collections were planted as a row of up to 5 plants. Trees were spaced at 3m within the row and 4m between rows. Meteorological data are given in the table as mean daily maximum and mean daily minimum temperature and mean total rainfall per month. Soil analyses were carried out in the Department of Forestry and Natural Resources, Edinburgh University, UK. The table shows the means of duplicate analyses of five samples taken from the genebank site and the forest surrounding the genebank. Soil data: pH 4.2 Org. Mat. (%) 9.4 N (Total, %) 0.3 P (Total, µg/g) 702 Ca (meq/100g) 5.19 Mg (meq/100g) 1.60 K (meq/100g) 0.48 Ca/Mg (ratio) 3.24 Ca+Mg/K (ratio) 14.1 Meteorological data: Temperature (°C) Rainfall (mm) Daily Max. Daily Min. January 30.9 21.2 116.2 February 29.1 20.6 272.3 March 29.8 21.2 260.0 April 29.9 21.3 327.5 May 29.9 21.0 287.9 June 27.9 20.4 308.1 July 28.1 20.0 254.1 August 29.3 20.3 262.5 September 30.3 20.5 282.4 October 30.6 21.1 257.1 November 30.8 21.1 361.2 December 31.0 21.4 154.8 Descriptors: The coding of the descriptors, and the methodology of recording, is based on the IBPGR standardised format, Engels et al., 1980 [ENG80A] and Engels, 1981 [ENG81A]. However, pod and bean data were recorded from a sample of 1-5 pods per collection, rather than the recommended sample of 10-20 pods due to limited availability of pods.

Flush Flower
Malaysia - BAL Plantations, 1991: Self Compatibility: self incompatible; Reference

Reference

Malaysia - BAL Plantations 1991
Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.
Notes

Malaysia - BAL Plantations Datafile: BAL2__DS MYS - BAL Plantations (update) Environment: Trials are planted on Table Estate, Tawau. Trial CA81 - planted August 1987 in 3x6 tree plots, 3 replicates randomised blocks, trees spaced at 3.8m x 2.4 m (1,096 trees/ha). Trial CA88 - similar design, planted August 1988. Soil type is fairly fertile young volcanic basalt. Average rainfall is c. 2100 mm/year well distributed, the driest month being February with about 150 mm. Descriptors: Cocoa butter analysis:- samples of a minimum of 2 kg were fermented in net bags placed in the fermentation heap and subsequently fast (i.e. continuously) dried. Determinations were carried out by MacRobertson Foods Pte Ltd. (a division of Cadbury-Schweppes Ltd.) in Singapore using a Nuclear Magnetic Resonance machine.

Yield Bean Butterfat Flower
Frances Bekele & Gillian Bidaisee, 2022: Ligule Colour: moderate anthocyanin; Stamen Colour: intense anthocyanin; Sepal Length: 7.71 mm; Ligule Width: 2.81 mm; Ovule Number: 55; Style Length: 2.24 mm; Pedicel Colour: reddish; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower

REFERENCES

Pound, F.J. (1938) Cacao and Witchbroom Disease of South America with notes on other species of Theobroma. Archives of Cocoa Research 1: pp. 26-71.

Zhang, D., Boccara, M., Motilal, L., Mischke S., Johnson, E.S., Butler, D.R., Bailey, B. & Meinhardt, L. (2009) Molecular characterization of an earliest cacao (Theobroma cacao L.) collection from Upper Amazon using microsatellite DNA markers. Tree Genetics & Genomes. doi:10.1007/s11295-009-0212-2

Motamayor, J.C., Lachenaud, P., da Silva e Mota, J.W., Loor, R., Kuhn, D.N., Brown, J.S. & Schnell, R.J. (2008) Geographic and Genetic Population Differentiation of the Amazonian Chocolate Tree (Theobroma cacao L). PLoS ONE 3(10): e3311. doi:10.1371/journal.pone.0003311

CEPEC (2012) Accession list for the CEPEC (Bahia, Brazil) collection. Dr. José Luis Pires, personal communication, February 2012.

Cote d'Ivoire - IRCC (1989) Liste des clones en collection. Unpublished Report, IRCC Bingerville, Cote d'Ivoire, Doc. No. 192. D. Clement, IRCC, Cote d'Ivoire. Personal Communication.

EET - Pichilingue (2012) Accessions planted in EET - Pichilingue, Ecuador. Freddy Amores and Ignacio Sotomayor C, personal communication, February 2012.

Malaysia - ARC Tuaran (1990) List of clones held in the Dept. of Agriculture's collection, at Sabah, Tuaran, Malaysia. E.B. Tay, Malaysian Cocoa Board, Malaysia. Personal Communication.

Togo - IRCC Togo (1991) Collection des clones de cacaoyers presents au Togo. Unpublished report, IRCC Togo, p.1.

CRU (2011) ICG,T accessions. CRU website.

Iwaro, D. (1998) Evalation of accessions for Black Pod. Personal Communication 30th October 1998. Data sent by email from Dr. Iwaro.

Lockwood, G. (1986) Report on cocoa breeding & seed production. Unpublished report for the Government of Ghana, Cocoa Research Institute of Ghana, CDC (UK.). pp. 23.

Lam, K.S. (1992) Performance of imported clones in Sime Darby Plantations. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.

Subali, S. (1992) MARDI's results on performance of imported primary cocoa clones. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.

Malaysia - Applied Agriculture (1992) Data from Applied Agriculture Research Unit (AARU). In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October 1992, Kuala Lumpur, Malaysia: pp. 4.

Malaysia - BAL Plantations (1991) Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.

Frances Bekele & Gillian Bidaisee (2022) Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.

Bartley, B.G.D. (1970) A preliminary catalogue of the cacao germplasm held at the University of the West Indies, Trinidad. University of the West Indies, St. Augustine, Trinidad.

Palaniappan, S. & Shuhaimi Shamsuddin (1992) Early evaluation of some cocoa germplasm at Pusut Perkhidmatan Pertanian Tun Razak. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.

Brazil - CEPEC (1987) List of clones in the CEPEC, Bahia, Brazil collection. Computer printout from CEPEC. Supplied by B.G.D. Bartley, CEPEC, Brazil.

Allen, J.B. (1988) Geographical variation and population biology in wild Theobroma cacao. PhD. Thesis, University of Edinburgh, U.K. pp. 197.

Trinidad - ICGT (1992) Leaf descriptors. Unpublished data supplied on computer diskette by International Cocoa Genebank, Trinidad, University of the West Indies, Trinidad.