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ICS 46 Group Name Theobroma cacao

Lockwood, G. & Gyamfi, M.M.O. (1979)
The CRIG cocoa germplasm collection with notes on codes used in the breeding programme at Tafo and elsewhere. Technical Bulletin, No. 10, Cocoa Research Institute, Tafo, Ghana. pp. 62.

Derivation: Imperial College Selections.
Location: ICTA, Trinidad and Tobago.

Notes: Lockwood & Gyamfi, 1979 [LOC79A] Selections from farms in Trinidad (1-100) and from Criollo progenies (101-107).

Cope, 1949a [COP49A]. One hundred selections were made as a result of a survey started in 1930 by Pound.

Bartley, 1993a [BAR93A]. Numbers 1 to 100 were selected from all over Trinidad, numbers 101 to 107 were Criollo trees selected from a farm but all have been lost and numbers greater than 111 are selections made in an experimental programme.

Bartley, 1996a [BAR96A]. The ICS material includes several clones which resulted from recent (at that time) introductions. These include Criollos from Nicaragua (ICS 60 and its relatives) or Venezuela. Other ICS clones represent "Amazonian" types originally introduced from the Orinoco. It is likely that several introductions were involved in creating the diversity that existed in Trinidad when Pound started the selection work.

Bartley, 1997b [BAR97B] From ICS 113 onwards these clones were used as parents in the AX / MX series. The same cross be known by an equivalent C [TTO] name.

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Synonyms: I S C 46

RECOMMENDED PLANTING MATERIAL Disclaimer

Recommended Planting Material

Disclaimer
Recommendations for planting materials are made by the referenced authors for specific regions and growing conditions. They are provided here as a guide and historic record only. To confirm the most appropriate planting material for your needs and its availability, please contact your local cocoa authority.

Details on clones that are available for distribution and guidelines on the safe movement of cacao germplasm can be obtained from the International Cocoa Quarantine Centre, Reading

SANGARE Abdourahamane et al., 2009: Recommended seed parent in:
Cote d'Ivoire; Parent of Hybrid: 'H 720'; Recommended by: Centre National de Recherche Agronomique (CNRA), Cote d'Ivoire; Reference

Reference

SANGARE Abdourahamane, KOFFI Edmond, AKAMOU Fataye & FALL Cheikh Alassane 2009
Rapport national sur l'état des ressources phytogénétiques pour l'alimentation et l'agriculture: République de Côte d’Ivoire. Ministère de l’agriculture
Notes

Additional information from Tahi et al. 2012

Recommended material
Efombagn Mousseni, Ives Bruno et al., 2009: Recommended seed parent in:
Cameroon; Female Parent in Seed Garden with: SNK 10 | SNK 13 | SNK 48 | SNK 64 | SNK 109 | ICS 43 | ICS 60 | T 79 /467 [POS] | SNK 12 | SNK 37 | SNK 450; Year: pre-1980; Reference

Reference

Efombagn Mousseni, Ives Bruno, Olivier Sounigo, Albertus B Eskes, Juan-Carlos Motamayor, Maria J Manzanares-Dauleux, Raymond J Schnell & Salomon Nyassé 2009
Parentage Analysis and Outcrossing Patterns in Cacao (Theobroma Cacao L.) Farms in Cameroon. Heredity 103 (1): 46–53
DOI: 10.1038/hdy.2009.30

Recommended material

PEDIGREE DETAILS

Mother to:: GEBP 303 /B-M [ADI] Frances Bekele, 2019

Reference

Frances Bekele 2019
Pedigree data for the GEBP clones supplied to ICQC,R (UK) by the Cocoa Research Centre (CRC), Trinidad & Tobago. Data supplied as an Excel spreadsheet by Frances Bekele, April 2019.; SI 46 /2 Trinidad - CRU, 2002

Reference

Trinidad - CRU 2002
ICG,T accessions database, reply to queries regarding names new to ICGD. Dr. David Butler, ICG,T Trinidad. Personal Communication. 12 September 2002.
Notes

Trinidad - International Cocoa Genebank, CRU, University of the West Indies Datafile: PROGENY_
Father to:: AHT 19 Nya Ngatchou, J., 1978

Reference

Nya Ngatchou, J. 1978
Etat d'avancement des travaux de génétique et d'amelioration du cacaoyer au Cameroun. In: Proceedings of the 7th International Cocoa Conference, Douala, Cameroun, 1979. Cocoa Producers' Alliance, Lagos, Nigeria. pp. 507-511.
Notes

Cameroon - IRAD Datafile: PROGENY_; H 720 [CIV] SANGARE Abdourahamane et al., 2009

Reference

SANGARE Abdourahamane, KOFFI Edmond, AKAMOU Fataye & FALL Cheikh Alassane 2009
Rapport national sur l'état des ressources phytogénétiques pour l'alimentation et l'agriculture: République de Côte d’Ivoire. Ministère de l’agriculture
Notes

Additional information from Tahi et al. 2012

Recommended material; SI 46 /2 Trinidad - CRU, 2002

Reference

Trinidad - CRU 2002
ICG,T accessions database, reply to queries regarding names new to ICGD. Dr. David Butler, ICG,T Trinidad. Personal Communication. 12 September 2002.
Notes

Trinidad - International Cocoa Genebank, CRU, University of the West Indies Datafile: PROGENY_

HELD IN

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United Kingdom, International Cocoa Quarantine Centre, Reading (ICQC,R): Current List ICQC,R Website; • Accession: RUQ 938 ICQC,R; - Local Name: ICS 46; - Year of Accession: 1999
Cameroon, Institut de Recherche Agricole pour le Développement (IRAD): List received in 1985 Reference

Reference

Cameroon - IRAD Cameroun 1985
Collection Camerounaise de cacaoyers (situation en Juillet, 1985). Unpublished report, IRCC Cameroun. Supplied by A.B. Eskes, IRCC Montpellier.; • Local Name: ICS 46
Cote d'Ivoire, Centre National de Recherche Agronomique (CNRA): List received in 1989 Reference

Reference

Cote d'Ivoire - IRCC 1989
Liste des clones en collection. Unpublished Report, IRCC Bingerville, Cote d'Ivoire, Doc. No. 192. D. Clement, IRCC, Cote d'Ivoire. Personal Communication.
Notes

List of clones grouped as West African Amelonados, Lower Amazons, Venezuelan collections, French Guiana collections, Ecuadorian collections, local Trinitarios, other Trinitarios, upper Amazons and Criollos. Notes on origin, date and place of origin and nature of the material (cuttings, budwood, seed).; • Local Name: ICS 46; - Year of Accession: 1957
French Guiana, CIRAD (Centre de Coopération Internationale en Researche Agronomique pour le Développment): List received in 2012 Reference

Reference

Philippe Lachenaud 2012
F. Guiana database, CIRAD-CP. Data supplied to ICGD by email. Dr. Philippe Lachenaud, CIRAD Montpellier, France, January 2012.; • Local Name: ICS 46; - Year of Accession: 1990
Gabon, SOCAGAB (Société de Développement de la Caféiculture et la Cacaoculture): List received in 1992 Reference

Reference

Gabon - SOCAGAB 1992
Liste des clones de la collection du C.M.C.E. de Minkong (Oyem, Gabon). Ph. Lachenaud, CIRAD, Kourou, Guyane. Personal Communication.; • Local Name: ICS 46
Guyana, National Agricultural Research Institute (NARI): List received in 2008 Reference

Reference

Chesney, P.E. 2008
Guyana accessions. Personal communication, September 2008.; • Local Name: ICS 46; - Year of Accession: 1954
India, Central Plantation Crops Research Institute (CPCRI): List received in 2008 Reference

Reference

CPCRI 2008
Cocoa germplasm maintained at the Central Plantation Crops Research Institute (CPCRI) Regional Station, Vittal, India. Data supplied to ICGD by email. D. Balasimha, CPCRI Vittal, India, July 2008.; • Accession: VTLC 188; - Local Name: ICS-46
Malaysia, Malaysian Cocoa Board (MCB): List received in 1998 Reference

Reference

Malaysia - ARC Tuaran 1998
Accessions held in the Malaysian Cocoa Board collection, May 1998. K. Lamin, MCB, Malaysia, Personal Communication.; • Local Name: ICS 46
Trinidad and Tobago, International Cocoa Genebank, Trinidad (ICG,T): List received in 2011 Reference

Reference

CRU 2011
ICG,T accessions. CRU website.
Notes

Website visited 12th April 2011

Link Unavailable [http://sta.uwi.edu/cru/icgt-accessions.asp]; • Local Name: ICS 46

DISEASE

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Phytophthora: Resistant; Butler, D.R. et al., 1998

Reference

Butler, D.R., Iwaro, A.D., Thevenin, J.-M., Bekele, F., Sounigo, O., Christopher, Y. & Mooledhar, V. 1998
Individual Institute Reports - Trinidad & Tobago, CRU. Report on the IPGRI project: Cocoa Germplasm Utilisation and Conservation, A Global Approach. Appendix to General Progress Report of Year 1.
Notes

Trinidad - ICG,T, University of the West Indies Datafile: DIS_PHDS - Disease reaction to Phytophthora DIS_WBDS - Disease reaction to Witches' Broom DIS_VSDS - Disease reaction to VSD DIS_CEDS - Disease reaction to Ceratocystis Descriptors: General observations and main traits of genotypes Environment: No details Germplasm: Clones included in the CFC/ICCO/IPGRI Cocoa Germplasm Utilisation and Conservation project.

Disease; Moderate Resistance, sp. P. palmivora (Pod).; Note: Unripe, Attached and not wounded; Morera, J., 1993

Reference

Morera, J. 1993
Reaction of clones to Phytophthora palmivora. J. Morera, CATIE, Costa Rica. Personal Communication.
Notes

Costa Rica - CATIE Datafile: DIS_PHDS - Disease reaction to Phytophthora Descriptors: Resistance to Phytophthora Environment: Costa Rica, CATIE. No further details Germplasm: 305 clones of various origins

Disease; Moderate Resistance, 90.00 % Infection, sp. P. palmivora (Pod).; Note: Unripe, Attached and not wounded; Blaha, G. & Lotode, R., 1976

Reference

Blaha, G. & Lotode, R. 1976
Un critere primordial de selection du cacaoyer au Cameroun: la resistance a la pourriture brune des cabossees ( Phytophthora palmivora ). Cafe, Cacao, The 20(2): pp. 97-116.
Notes

Cameroon - IFCC Datafile: DIS_PHDS - Disease reaction to Phytophthora Decriptors: Resistance based on inoculation with zoospore suspension of P. palmivora Environment: IFCC, Cameroon Germplasm: 80 clones of trinitario and forestero type (SNK), 14 clones from Trinidad of criollo hybrids (ICS), five clones of Amazonian forestero (UPA) and hybrids produced at Tafo from recent introductions of IMC 60, IMC 67, NA 32, PA 7 [PER] and SCA 6.

Disease; Moderately Susceptible, sp. P. palmivora (Pod).; Note: Unripe, Unattached and not wounded; Iwaro, D., 1998

Reference

Iwaro, D. 1998
Evalation of accessions for Black Pod. Personal Communication 30th October 1998. Data sent by email from Dr. Iwaro.
Notes

Trinidad - CRU, University of the West Indies Datafile: DIS_PHDS - Disease reaction to Phytophthora Experimental conditions: Inoculation techinique used - spray method. Inoculum concentration: 100,000 zoospores/ml. Incubated for four days. Character Data: The following descriptions are used to segregate clones according to reaction: no visible symptoms = highly resistant to penetration, 1-5 localised lesions = resistant, 6-15 localised lesions = moderately resistant, >15 localised lesions = partially resistant/resistant to spread of lesion alone, 1-5 expanding lesions = partially resistant/resistant to penetration alone, 6-15 expanding lesions = moderately susceptible, >15 expanding lesions = susceptible, coalesced lesions = highly susceptible Disease - Phytophthora palmivora, P. megakarya (Local Names: Black Pod, Podredumbre negra, Pourriture Brune, Podridao parda).

Disease; Resistant, sp. P. palmivora (Pod).; Trinidad - CRU, 1994

Reference

Trinidad - CRU 1994
Cocoa Research Unit. Report for 1994. The University of the West Indies.
Notes

Trinidad - CRU, University of the West Indies Datafile: ICGTF_DS TTO - ICG,T Flower Characters ICGTFRDS TTO - ICG,T Fruit Characters ICGTL_DS TTO - ICG,T Leaf Characters DIS_WBDS - Disease reaction to Witches' Broom Germplasm: Flower descriptor data is presented for 620 clones. Fruit descriptor data is presented for 333 clones. The seed characters of 118 clones are partly described. Leaf descriptor data is presented for 229 clones. The germplasm is mainly of Peruvian, Ecuadorian and Colombian origin. Environment: Each clone is represented in the collection by up to 16 trees planted 1.8 m apart in a block design. Shading is provided by Erythrina sp. (spaced 60m apart) and bananas (spaced 4m apart). Fertilizer, N-P-K (15:10:5), is provided twice a year at 250g per tree. Cyclic irrigation is performed from February to May (10 l of water per plant/week). Soil type group -33 A 1 Soil Series - Cunupia fine sandy clay with restricted internal drainage Lithology - silty clay alluvium Drainage - imperfect Slope category - 0°-2° Meterological data are given in this table as mean maximum, mean minimum and overall mean temperature and mean total rainfall per month over the period 1961-1990. Month Temp. (°C) Rainfall Max Min Mean (mm) January 30.1 20.8 24.6 71.2 February 30.2 20.8 25.0 43.3 March 31.2 21.4 25.7 36.6 April 31.8 22.3 26.5 51.1 May 31.6 23.3 26.9 116.5 June 30.6 23.3 26.4 252.1 July 30.8 23.0 26.2 266.3 August 31.2 22.9 26.4 250.1 September 31.5 22.8 26.5 202.5 October 31.4 22.82 6.41 99.3 November 30.8 22.4 25.9 227.8 December 30.2 21.5 25.2 155.6 Descriptors: The character selection is based on the IBPGR list (IBPGR, 1981 [IBP81A]). Where descriptor states present real measurements, e.g. pod length in cm, the data are given as the mean of all the measurements ± the standard deviation. For some descriptors, such as Leaf, flush colour, the phenotypic expression of the character is coded on a scale from 0 to 9. On this scale '0' represents the lowest or weakest expression of a characteristic and '9' represents the highest or strongest. Leaf and Flower Characteristics p.300-307 Morphological Characteristics of Flowers, Fruit and Leaves- ICGT, Trinidad, Disease - Crinipellis perniciosa (Marasmius perniciosa) (local names: Witches' Broom Disease, Escoba de bruja, Balai de sorciere, Vassoura de bruxa).

Disease; Tolerant, sp. P. palmivora (Pod).; Enriquez, G. & Soria, V.J., 1977

Reference

Enriquez, G. & Soria, V.J. 1977
Mejoramiento genético para resistencia a cinco enfermedades del cacao. In: Proceedings of the 6th International Cocoa Research Conference, Caracas, Venezuela, 1977. Cocoa Producers' Alliance, Lagos, Nigeria. pp. 45-56.
Notes

Various data sources Review of disease data complied at Costa Rica - CATIE from sources including: Costa Rica - CATIE, Trinidad - CRU, University of the West Indies, Ghana - WACRI, Ecuador - Pichilingue. Datafile: DIS_MODS - Disease reaction to Monilia Pod Rot DIS___DS - Disease reaction to stress and disease DIS_PHDS - Disease reaction to Phytophthora DIS_SSDS - Disease reaction to CSSV DIS_WBDS - Disease reaction to Witches' Broom DIS_CEDS - Disease reaction to Ceratocystis Descriptors: Information on the type of disease/stress and the reaction of the clone.

Disease
Witches Broom: Susceptible (Field Reaction).; Butler, D.R. et al., 1998

Reference

Butler, D.R., Iwaro, A.D., Thevenin, J.-M., Bekele, F., Sounigo, O., Christopher, Y. & Mooledhar, V. 1998
Individual Institute Reports - Trinidad & Tobago, CRU. Report on the IPGRI project: Cocoa Germplasm Utilisation and Conservation, A Global Approach. Appendix to General Progress Report of Year 1.
Notes

Trinidad - ICG,T, University of the West Indies Datafile: DIS_PHDS - Disease reaction to Phytophthora DIS_WBDS - Disease reaction to Witches' Broom DIS_VSDS - Disease reaction to VSD DIS_CEDS - Disease reaction to Ceratocystis Descriptors: General observations and main traits of genotypes Environment: No details Germplasm: Clones included in the CFC/ICCO/IPGRI Cocoa Germplasm Utilisation and Conservation project.

Disease; Tolerant (Pod).; Trinidad - CRU, 1994

Reference

Trinidad - CRU 1994
Cocoa Research Unit. Report for 1994. The University of the West Indies.
Notes

Trinidad - CRU, University of the West Indies Datafile: ICGTF_DS TTO - ICG,T Flower Characters ICGTFRDS TTO - ICG,T Fruit Characters ICGTL_DS TTO - ICG,T Leaf Characters DIS_WBDS - Disease reaction to Witches' Broom Germplasm: Flower descriptor data is presented for 620 clones. Fruit descriptor data is presented for 333 clones. The seed characters of 118 clones are partly described. Leaf descriptor data is presented for 229 clones. The germplasm is mainly of Peruvian, Ecuadorian and Colombian origin. Environment: Each clone is represented in the collection by up to 16 trees planted 1.8 m apart in a block design. Shading is provided by Erythrina sp. (spaced 60m apart) and bananas (spaced 4m apart). Fertilizer, N-P-K (15:10:5), is provided twice a year at 250g per tree. Cyclic irrigation is performed from February to May (10 l of water per plant/week). Soil type group -33 A 1 Soil Series - Cunupia fine sandy clay with restricted internal drainage Lithology - silty clay alluvium Drainage - imperfect Slope category - 0°-2° Meterological data are given in this table as mean maximum, mean minimum and overall mean temperature and mean total rainfall per month over the period 1961-1990. Month Temp. (°C) Rainfall Max Min Mean (mm) January 30.1 20.8 24.6 71.2 February 30.2 20.8 25.0 43.3 March 31.2 21.4 25.7 36.6 April 31.8 22.3 26.5 51.1 May 31.6 23.3 26.9 116.5 June 30.6 23.3 26.4 252.1 July 30.8 23.0 26.2 266.3 August 31.2 22.9 26.4 250.1 September 31.5 22.8 26.5 202.5 October 31.4 22.82 6.41 99.3 November 30.8 22.4 25.9 227.8 December 30.2 21.5 25.2 155.6 Descriptors: The character selection is based on the IBPGR list (IBPGR, 1981 [IBP81A]). Where descriptor states present real measurements, e.g. pod length in cm, the data are given as the mean of all the measurements ± the standard deviation. For some descriptors, such as Leaf, flush colour, the phenotypic expression of the character is coded on a scale from 0 to 9. On this scale '0' represents the lowest or weakest expression of a characteristic and '9' represents the highest or strongest. Leaf and Flower Characteristics p.300-307 Morphological Characteristics of Flowers, Fruit and Leaves- ICGT, Trinidad, Disease - Crinipellis perniciosa (Marasmius perniciosa) (local names: Witches' Broom Disease, Escoba de bruja, Balai de sorciere, Vassoura de bruxa).

Disease

PESTS

Mirids: Resistant; K. F. N’Guessan et al., 2003

Reference

K. F. N’Guessan, A. B. Eskes & Ph. Lachenaud 2003
Etude de la Resistance du Cacaoyer aux Mirides : Classification des Grands Groupes de Cacaoyers en Fonction de Leur Niveau de Resistance ou Sensibilite. In: Proceedings of the 14th International Cocoa Research Conference, Accra, Ghana (October 2003). Cocoa Producers' Alliance, Lagos, Nigeria.
Notes

Title: 'Study on the Resistance of Cocoa Trees to Mirids in Cote d’Ivoire: Classification of Major Cocoa Tree Groups by their Level of Resistance / Sensitivity'.

Pests

YIELD

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Malaysia - BAL Plantations, 1991: Seed Index: 66.6 (dried beans per 100g); Pod Index: 19.4 (pods per kg dried beans); Reference

Reference

Malaysia - BAL Plantations 1991
Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.
Notes

Malaysia - BAL Plantations Datafile: BAL2__DS MYS - BAL Plantations (update) Environment: Trials are planted on Table Estate, Tawau. Trial CA81 - planted August 1987 in 3x6 tree plots, 3 replicates randomised blocks, trees spaced at 3.8m x 2.4 m (1,096 trees/ha). Trial CA88 - similar design, planted August 1988. Soil type is fairly fertile young volcanic basalt. Average rainfall is c. 2100 mm/year well distributed, the driest month being February with about 150 mm. Descriptors: Cocoa butter analysis:- samples of a minimum of 2 kg were fermented in net bags placed in the fermentation heap and subsequently fast (i.e. continuously) dried. Determinations were carried out by MacRobertson Foods Pte Ltd. (a division of Cadbury-Schweppes Ltd.) in Singapore using a Nuclear Magnetic Resonance machine.

Yield Bean Butterfat Flower
Frances Bekele & Gillian Bidaisee, 2022: Pod Index: 22.6 (pods per kg dried beans); Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Costa Rica - CATIE, 1981: Productivity: low; Reference

Reference

Costa Rica - CATIE 1981
Genetic Resources of Cacao. A Catalogue of the CATIE Collection. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.
Notes

Costa Rica - CATIE Datafile: CATIE__DS CRI - CATIE 1 IMAGE_DS PROGENY_ Germplasm: Descriptor information is presented for 294 clones, 8 related species and on one hybrid. In addition, limited information on a further 119 clones is given. This germplasm is mainly of Latin American origin, including the United Fruit clones, UF, and genotypes from Mexico and Brazil. Environment: Each clone is represented in the collection by up to 10 trees planted in a single row. Spacing is usually 2m within the row and 3m between the rows, although parts of the collection are planted in a 2.2 x 2.2m grid. Fertilizer, N-P-K-Mg/Ca in the ratio 18:10:6:5, is applied four times a year at 150g per tree. Shade is provided in most of the collection by an almost complete closed cover of trees of Erythrina poepiggiana and Inga spp. spaced at 12 x 12m. Character data: Seed dry weight: sample is dried for 1.5 h at 130°C and cooled in a desiccator. Seed length, seed width, seed thickness: average of 5 seeds from 20 fruits. Fruit, wall thickness at ridge: fruits are transversally sectioned and the soft endocarp tissue removed. Fruit, ridge pair separation: mean ratio of the distance between ridges within a pair and the distances between the ridges of two different pairs. Leaf, flush colour: measured when leaves are 5cm long. Producitvity: values are given as: 1=very low - c. 40 pods harvested over 3 years; 3=low, 5=intermediate - c. 40 pods obtained from 3-4 harvests over 1 year; 7=high, 9=very high - c. 40 pods harvested at one time. Pod outline drawing for each accession.

Yield Bean Fruit Flush Flower

BEAN

Show Details Hide Details

Malaysia - BAL Plantations, 1991: Number: 34.2; Total Dry Weight: 51.6 g; Dry Weight: 1.50 g; Note: 27 plants sampled; Reference

Reference

Malaysia - BAL Plantations 1991
Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.
Notes

Malaysia - BAL Plantations Datafile: BAL2__DS MYS - BAL Plantations (update) Environment: Trials are planted on Table Estate, Tawau. Trial CA81 - planted August 1987 in 3x6 tree plots, 3 replicates randomised blocks, trees spaced at 3.8m x 2.4 m (1,096 trees/ha). Trial CA88 - similar design, planted August 1988. Soil type is fairly fertile young volcanic basalt. Average rainfall is c. 2100 mm/year well distributed, the driest month being February with about 150 mm. Descriptors: Cocoa butter analysis:- samples of a minimum of 2 kg were fermented in net bags placed in the fermentation heap and subsequently fast (i.e. continuously) dried. Determinations were carried out by MacRobertson Foods Pte Ltd. (a division of Cadbury-Schweppes Ltd.) in Singapore using a Nuclear Magnetic Resonance machine.

Yield Bean Butterfat Flower
Bartley, B.G.D., 1970: Size: large; Reference

Reference

Bartley, B.G.D. 1970
A preliminary catalogue of the cacao germplasm held at the University of the West Indies, Trinidad. University of the West Indies, St. Augustine, Trinidad.
Notes

Trinidad - University of the West Indies Datafile: ICGT2_DS TTO - UWI Cacao Collection Germplasm: Passport descriptors for approximately 1000 accessions of mainly South American and Trinidadian origin. Environment: No details. Descriptors: Six descriptors for fruit, seed and compatability characteristics. Character Data: Combinations used FXC = forastero x criollo, TX = trinitario cross

Bean Fruit Flower
Frances Bekele & Gillian Bidaisee, 2022: Colour: purple; Shape: elliptical; Number: 32.0; Total Wet Weight: 68.0 g; Cotyledon Dry Weight: 1.38 g; Cotyledon Length: 25.2 mm; Cotyledon Width: 14.7 mm; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Palaniappan, S. & Shuhaimi Shamsuddin, 1992: Number: 36.4; Dry Weight: 1.58 g; Reference

Reference

Palaniappan, S. & Shuhaimi Shamsuddin 1992
Early evaluation of some cocoa germplasm at Pusut Perkhidmatan Pertanian Tun Razak. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Notes

Malaysia - FELDA, Pusat Perkhidmatan Pertanian Tun Razak, Peninsula Malaysia Datafile: FELDA_DC MYS - Felda PPPTR Germplasm: Agronomic and morphological characteristics of some of the clones held at Pusat Perkhidmatan Pertanian Tun Razak. The clones were obtained from the Malaysian Cocoa Growers Council Joint Importation Scheme, MARDI and the Department of Agriculture, Peninsular Malaysia. Environment: The clones are planted in single plots of 18 plants/clone (6x3) at Pusat Perkhidmatan Pertanian Tun Razak, Sg. Tekam in inland Peninsular Malaysia. The clones were planted as they were obtained and therefore are of varying age. Character data: Seed dry weight: corrected to 6% moisture after oven drying at 105'C for 24 hours.

Bean Fruit

BUTTERFAT

Malaysia - BAL Plantations, 1991: Butterfat Content: 54.90 %; Maximum Butterfat Content: 56.32 %; Minimum Butterfat Content: 54.03 %; Note: 7 plants sampled; Reference

Reference

Malaysia - BAL Plantations 1991
Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.
Notes

Malaysia - BAL Plantations Datafile: BAL2__DS MYS - BAL Plantations (update) Environment: Trials are planted on Table Estate, Tawau. Trial CA81 - planted August 1987 in 3x6 tree plots, 3 replicates randomised blocks, trees spaced at 3.8m x 2.4 m (1,096 trees/ha). Trial CA88 - similar design, planted August 1988. Soil type is fairly fertile young volcanic basalt. Average rainfall is c. 2100 mm/year well distributed, the driest month being February with about 150 mm. Descriptors: Cocoa butter analysis:- samples of a minimum of 2 kg were fermented in net bags placed in the fermentation heap and subsequently fast (i.e. continuously) dried. Determinations were carried out by MacRobertson Foods Pte Ltd. (a division of Cadbury-Schweppes Ltd.) in Singapore using a Nuclear Magnetic Resonance machine.

Yield Bean Butterfat Flower
Frances Bekele & Gillian Bidaisee, 2022: Testa: 4.80 %; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower

FRUIT

Show Details Hide Details

Bartley, B.G.D., 1970: Size: large; Reference

Reference

Bartley, B.G.D. 1970
A preliminary catalogue of the cacao germplasm held at the University of the West Indies, Trinidad. University of the West Indies, St. Augustine, Trinidad.
Notes

Trinidad - University of the West Indies Datafile: ICGT2_DS TTO - UWI Cacao Collection Germplasm: Passport descriptors for approximately 1000 accessions of mainly South American and Trinidadian origin. Environment: No details. Descriptors: Six descriptors for fruit, seed and compatability characteristics. Character Data: Combinations used FXC = forastero x criollo, TX = trinitario cross

Bean Fruit Flower
Frances Bekele & Gillian Bidaisee, 2022: Shape: oblate; Basal Constriction: absent; Apex Form: attenuate; Rugosity: intense; Length: 171.0 mm; Breadth: 84.0 mm; Colour: anthocyanin absent; Hardness: hard; Note: Hardness > 2.0 Mpa; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Costa Rica - CATIE, 1981: Basal Constriction: absent; Apex Form: acute; Rugosity: slight; Colour: slight / absent anthocyanin (ridge/furrow); Pods Per Tree: 3; Reference

Reference

Costa Rica - CATIE 1981
Genetic Resources of Cacao. A Catalogue of the CATIE Collection. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.
Notes

Costa Rica - CATIE Datafile: CATIE__DS CRI - CATIE 1 IMAGE_DS PROGENY_ Germplasm: Descriptor information is presented for 294 clones, 8 related species and on one hybrid. In addition, limited information on a further 119 clones is given. This germplasm is mainly of Latin American origin, including the United Fruit clones, UF, and genotypes from Mexico and Brazil. Environment: Each clone is represented in the collection by up to 10 trees planted in a single row. Spacing is usually 2m within the row and 3m between the rows, although parts of the collection are planted in a 2.2 x 2.2m grid. Fertilizer, N-P-K-Mg/Ca in the ratio 18:10:6:5, is applied four times a year at 150g per tree. Shade is provided in most of the collection by an almost complete closed cover of trees of Erythrina poepiggiana and Inga spp. spaced at 12 x 12m. Character data: Seed dry weight: sample is dried for 1.5 h at 130°C and cooled in a desiccator. Seed length, seed width, seed thickness: average of 5 seeds from 20 fruits. Fruit, wall thickness at ridge: fruits are transversally sectioned and the soft endocarp tissue removed. Fruit, ridge pair separation: mean ratio of the distance between ridges within a pair and the distances between the ridges of two different pairs. Leaf, flush colour: measured when leaves are 5cm long. Producitvity: values are given as: 1=very low - c. 40 pods harvested over 3 years; 3=low, 5=intermediate - c. 40 pods obtained from 3-4 harvests over 1 year; 7=high, 9=very high - c. 40 pods harvested at one time. Pod outline drawing for each accession.

Yield Bean Fruit Flush Flower
Morera, J. et al., 1991: Shape: calabacillo; Colour: green; Reference

Reference

Morera, J., Paredes, A. & Mora, A. 1991
Germoplasma de cacao en el CATIE entre 1947 y 1991, Programa II: Generacion y transferencia de tecnologia. IICA, San Jose, Costa Rica, pp. 49.
Notes

Costa Rica - CATIE Datafile: DIS_MODS - Disease reaction to Monilia Pod Rot CATIE3DS CRI - CATIE 3 DIS_PHDS - Disease reaction to Phytophthora DIS_CEDS - Disease reaction to Ceratocystis PROGENY_ Germplasm: Descriptions are given for 559 clones. More information on 44 of them is given in the Table CATIE, Costa Rica (1). Environment: Details are given in the Table CATIE, Costa Rica (1). Descriptors: Population type, self-compatibility, resistance to disease, pod shape and colour are described. Population types - CATIE, Costa Rica (3). Disease - Phytophthora palmivora, P. megakarya (Local Names: Black Pod, Podredumbre negra, Pourriture Brune, Podridao parda), Moniliophthora / Monilia roreri (local names: Watery Pod Rot, Moniliasis, Moniliose), Ceratocystis fimbriata (local names: Ceratocystis Wilt, Mal de Machete, Mort Subite).

Disease Fruit Flower
Palaniappan, S. & Shuhaimi Shamsuddin, 1992: Length: 165.7 mm; Breadth: 82.8 mm; Weight: 498.5 g; Length Breadth Ratio: 2.00; Reference

Reference

Palaniappan, S. & Shuhaimi Shamsuddin 1992
Early evaluation of some cocoa germplasm at Pusut Perkhidmatan Pertanian Tun Razak. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Notes

Malaysia - FELDA, Pusat Perkhidmatan Pertanian Tun Razak, Peninsula Malaysia Datafile: FELDA_DC MYS - Felda PPPTR Germplasm: Agronomic and morphological characteristics of some of the clones held at Pusat Perkhidmatan Pertanian Tun Razak. The clones were obtained from the Malaysian Cocoa Growers Council Joint Importation Scheme, MARDI and the Department of Agriculture, Peninsular Malaysia. Environment: The clones are planted in single plots of 18 plants/clone (6x3) at Pusat Perkhidmatan Pertanian Tun Razak, Sg. Tekam in inland Peninsular Malaysia. The clones were planted as they were obtained and therefore are of varying age. Character data: Seed dry weight: corrected to 6% moisture after oven drying at 105'C for 24 hours.

Bean Fruit

FLUSH

Frances Bekele & Gillian Bidaisee, 2022: Colour: moderate anthocyanin; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Costa Rica - CATIE, 1981: Colour: olive yellow; Reference

Reference

Costa Rica - CATIE 1981
Genetic Resources of Cacao. A Catalogue of the CATIE Collection. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.
Notes

Costa Rica - CATIE Datafile: CATIE__DS CRI - CATIE 1 IMAGE_DS PROGENY_ Germplasm: Descriptor information is presented for 294 clones, 8 related species and on one hybrid. In addition, limited information on a further 119 clones is given. This germplasm is mainly of Latin American origin, including the United Fruit clones, UF, and genotypes from Mexico and Brazil. Environment: Each clone is represented in the collection by up to 10 trees planted in a single row. Spacing is usually 2m within the row and 3m between the rows, although parts of the collection are planted in a 2.2 x 2.2m grid. Fertilizer, N-P-K-Mg/Ca in the ratio 18:10:6:5, is applied four times a year at 150g per tree. Shade is provided in most of the collection by an almost complete closed cover of trees of Erythrina poepiggiana and Inga spp. spaced at 12 x 12m. Character data: Seed dry weight: sample is dried for 1.5 h at 130°C and cooled in a desiccator. Seed length, seed width, seed thickness: average of 5 seeds from 20 fruits. Fruit, wall thickness at ridge: fruits are transversally sectioned and the soft endocarp tissue removed. Fruit, ridge pair separation: mean ratio of the distance between ridges within a pair and the distances between the ridges of two different pairs. Leaf, flush colour: measured when leaves are 5cm long. Producitvity: values are given as: 1=very low - c. 40 pods harvested over 3 years; 3=low, 5=intermediate - c. 40 pods obtained from 3-4 harvests over 1 year; 7=high, 9=very high - c. 40 pods harvested at one time. Pod outline drawing for each accession.

Yield Bean Fruit Flush Flower

LEAF

Trinidad - ICGT, 1992: Distance from Base to Widest Part: 139 mm; Length: 293.0 mm; Width: 102.0 mm; Length to Width Ratio: 2.40; Basal Angle: 110.8 °; Apical Angle: 86.6 °; Petiole Length: 28 mm; Pulvinus Length: 21 mm; Reference

Reference

Trinidad - ICGT 1992
Leaf descriptors. Unpublished data supplied on computer diskette by International Cocoa Genebank, Trinidad, University of the West Indies, Trinidad.
Notes

Trinidad - International Cocoa Genebank, CRU, University of the West Indies.
Flower descriptor data is presented for 620 clones, fruit descriptor data of 333 and seed characters of 118 clones are partly described. Leaf descriptor data is presented for 229 clones. The germplasm is mainly of Peruvian, Ecuadorian and Colombian origin. Environment: Each clone is represented in the collection by up to 16 trees planted 1.8 m apart in a block design. Shading is provided by Erythrina sp. (spaced 60m apart) and bananas (spaced 4m apart). Fertilizer, N-P-K (15:10:5), is provided twice a year at 250g per tree. Cyclic irrigation is performed from February to May (10 l of water per plant/week). Soil type group -33 A 1 Soil Series - Cunupia fine sandy clay with restricted internal drainage Lithology - silty clay alluvium Drainage - imperfect Slope category - 0°-2° Meterological data are given in this table as mean maximum, mean minimum and overall mean temperature and mean total rainfall per month over the period 1961-1990. Month Temp. (°C) Rainfall Max Min Mean (mm) January 30.1 20.8 24.6 71.2 February 30.2 20.8 25.0 43.3 March 31.2 21.4 25.7 36.6 April 31.8 22.3 26.5 51.1 May 31.6 23.3 26.9 116.5 June 30.6 23.3 26.4 252.1 July 30.8 23.0 26.2 266.3 August 31.2 22.9 26.4 250.1 September 31.5 22.8 26.5 202.5 October 31.4 22.82 6.41 99.3 November 30.8 22.4 25.9 227.8 December 30.2 21.5 25.2 155.6 Descriptors: The character selection is based on the IBPGR list (IBPGR, 1981 [IBP81A]). Where descriptor states present real measurements, e.g. pod length in cm, the data are given as the mean of all the measurements ± the standard deviation. For some descriptors, such as Leaf, flush colour, the phenotypic expression of the character is coded on a scale from 0 to 9. On this scale '0' represents the lowest or weakest expression of a characteristic and '9' represents the highest or strongest. Leaf and Flower Characteristics p.300-307

Leaf

FLOWER

Show Details Hide Details

Malaysia - BAL Plantations, 1991: Self Compatibility: self incompatible; Reference

Reference

Malaysia - BAL Plantations 1991
Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.
Notes

Malaysia - BAL Plantations Datafile: BAL2__DS MYS - BAL Plantations (update) Environment: Trials are planted on Table Estate, Tawau. Trial CA81 - planted August 1987 in 3x6 tree plots, 3 replicates randomised blocks, trees spaced at 3.8m x 2.4 m (1,096 trees/ha). Trial CA88 - similar design, planted August 1988. Soil type is fairly fertile young volcanic basalt. Average rainfall is c. 2100 mm/year well distributed, the driest month being February with about 150 mm. Descriptors: Cocoa butter analysis:- samples of a minimum of 2 kg were fermented in net bags placed in the fermentation heap and subsequently fast (i.e. continuously) dried. Determinations were carried out by MacRobertson Foods Pte Ltd. (a division of Cadbury-Schweppes Ltd.) in Singapore using a Nuclear Magnetic Resonance machine.

Yield Bean Butterfat Flower
Bartley, B.G.D., 1970: Self Compatibility: self incompatible; Reference

Reference

Bartley, B.G.D. 1970
A preliminary catalogue of the cacao germplasm held at the University of the West Indies, Trinidad. University of the West Indies, St. Augustine, Trinidad.
Notes

Trinidad - University of the West Indies Datafile: ICGT2_DS TTO - UWI Cacao Collection Germplasm: Passport descriptors for approximately 1000 accessions of mainly South American and Trinidadian origin. Environment: No details. Descriptors: Six descriptors for fruit, seed and compatability characteristics. Character Data: Combinations used FXC = forastero x criollo, TX = trinitario cross

Bean Fruit Flower
Frances Bekele & Gillian Bidaisee, 2022: Ligule Colour: anthocyanin absent; Stamen Colour: anthocyanin absent; Sepal Length: 8.50 mm; Ligule Width: 2.59 mm; Ovule Number: 38; Style Length: 1.85 mm; Pedicel Colour: red; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Morera, J. et al., 1991: Self Compatibility: self compatible; Reference

Reference

Morera, J., Paredes, A. & Mora, A. 1991
Germoplasma de cacao en el CATIE entre 1947 y 1991, Programa II: Generacion y transferencia de tecnologia. IICA, San Jose, Costa Rica, pp. 49.
Notes

Costa Rica - CATIE Datafile: DIS_MODS - Disease reaction to Monilia Pod Rot CATIE3DS CRI - CATIE 3 DIS_PHDS - Disease reaction to Phytophthora DIS_CEDS - Disease reaction to Ceratocystis PROGENY_ Germplasm: Descriptions are given for 559 clones. More information on 44 of them is given in the Table CATIE, Costa Rica (1). Environment: Details are given in the Table CATIE, Costa Rica (1). Descriptors: Population type, self-compatibility, resistance to disease, pod shape and colour are described. Population types - CATIE, Costa Rica (3). Disease - Phytophthora palmivora, P. megakarya (Local Names: Black Pod, Podredumbre negra, Pourriture Brune, Podridao parda), Moniliophthora / Monilia roreri (local names: Watery Pod Rot, Moniliasis, Moniliose), Ceratocystis fimbriata (local names: Ceratocystis Wilt, Mal de Machete, Mort Subite).

Disease Fruit Flower

REFERENCES

Lockwood, G. & Gyamfi, M.M.O. (1979) The CRIG cocoa germplasm collection with notes on codes used in the breeding programme at Tafo and elsewhere. Technical Bulletin, No. 10, Cocoa Research Institute, Tafo, Ghana. pp. 62.

SANGARE Abdourahamane, KOFFI Edmond, AKAMOU Fataye & FALL Cheikh Alassane (2009) Rapport national sur l'état des ressources phytogénétiques pour l'alimentation et l'agriculture: République de Côte d’Ivoire. Ministère de l’agriculture

Efombagn Mousseni, Ives Bruno, Olivier Sounigo, Albertus B Eskes, Juan-Carlos Motamayor, Maria J Manzanares-Dauleux, Raymond J Schnell & Salomon Nyassé (2009) Parentage Analysis and Outcrossing Patterns in Cacao (Theobroma Cacao L.) Farms in Cameroon. Heredity 103 (1): 46–53

Frances Bekele (2019) Pedigree data for the GEBP clones supplied to ICQC,R (UK) by the Cocoa Research Centre (CRC), Trinidad & Tobago. Data supplied as an Excel spreadsheet by Frances Bekele, April 2019.

Trinidad - CRU (2002) ICG,T accessions database, reply to queries regarding names new to ICGD. Dr. David Butler, ICG,T Trinidad. Personal Communication. 12 September 2002.

Nya Ngatchou, J. (1978) Etat d'avancement des travaux de génétique et d'amelioration du cacaoyer au Cameroun. In: Proceedings of the 7th International Cocoa Conference, Douala, Cameroun, 1979. Cocoa Producers' Alliance, Lagos, Nigeria. pp. 507-511.

Cameroon - IRAD Cameroun (1985) Collection Camerounaise de cacaoyers (situation en Juillet, 1985). Unpublished report, IRCC Cameroun. Supplied by A.B. Eskes, IRCC Montpellier.

Cote d'Ivoire - IRCC (1989) Liste des clones en collection. Unpublished Report, IRCC Bingerville, Cote d'Ivoire, Doc. No. 192. D. Clement, IRCC, Cote d'Ivoire. Personal Communication.

Philippe Lachenaud (2012) F. Guiana database, CIRAD-CP. Data supplied to ICGD by email. Dr. Philippe Lachenaud, CIRAD Montpellier, France, January 2012.

Gabon - SOCAGAB (1992) Liste des clones de la collection du C.M.C.E. de Minkong (Oyem, Gabon). Ph. Lachenaud, CIRAD, Kourou, Guyane. Personal Communication.

Chesney, P.E. (2008) Guyana accessions. Personal communication, September 2008.

CPCRI (2008) Cocoa germplasm maintained at the Central Plantation Crops Research Institute (CPCRI) Regional Station, Vittal, India. Data supplied to ICGD by email. D. Balasimha, CPCRI Vittal, India, July 2008.

Malaysia - ARC Tuaran (1998) Accessions held in the Malaysian Cocoa Board collection, May 1998. K. Lamin, MCB, Malaysia, Personal Communication.

CRU (2011) ICG,T accessions. CRU website.

Butler, D.R., Iwaro, A.D., Thevenin, J.-M., Bekele, F., Sounigo, O., Christopher, Y. & Mooledhar, V. (1998) Individual Institute Reports - Trinidad & Tobago, CRU. Report on the IPGRI project: Cocoa Germplasm Utilisation and Conservation, A Global Approach. Appendix to General Progress Report of Year 1.

Morera, J. (1993) Reaction of clones to Phytophthora palmivora. J. Morera, CATIE, Costa Rica. Personal Communication.

Blaha, G. & Lotode, R. (1976) Un critere primordial de selection du cacaoyer au Cameroun: la resistance a la pourriture brune des cabossees ( Phytophthora palmivora ). Cafe, Cacao, The 20(2): pp. 97-116.

Iwaro, D. (1998) Evalation of accessions for Black Pod. Personal Communication 30th October 1998. Data sent by email from Dr. Iwaro.

Trinidad - CRU (1994) Cocoa Research Unit. Report for 1994. The University of the West Indies.

Enriquez, G. & Soria, V.J. (1977) Mejoramiento genético para resistencia a cinco enfermedades del cacao. In: Proceedings of the 6th International Cocoa Research Conference, Caracas, Venezuela, 1977. Cocoa Producers' Alliance, Lagos, Nigeria. pp. 45-56.

K. F. N’Guessan, A. B. Eskes & Ph. Lachenaud (2003) Etude de la Resistance du Cacaoyer aux Mirides : Classification des Grands Groupes de Cacaoyers en Fonction de Leur Niveau de Resistance ou Sensibilite. In: Proceedings of the 14th International Cocoa Research Conference, Accra, Ghana (October 2003). Cocoa Producers' Alliance, Lagos, Nigeria.

Malaysia - BAL Plantations (1991) Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.

Frances Bekele & Gillian Bidaisee (2022) Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.

Costa Rica - CATIE (1981) Genetic Resources of Cacao. A Catalogue of the CATIE Collection. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.

Bartley, B.G.D. (1970) A preliminary catalogue of the cacao germplasm held at the University of the West Indies, Trinidad. University of the West Indies, St. Augustine, Trinidad.

Palaniappan, S. & Shuhaimi Shamsuddin (1992) Early evaluation of some cocoa germplasm at Pusut Perkhidmatan Pertanian Tun Razak. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.

Morera, J., Paredes, A. & Mora, A. (1991) Germoplasma de cacao en el CATIE entre 1947 y 1991, Programa II: Generacion y transferencia de tecnologia. IICA, San Jose, Costa Rica, pp. 49.

Trinidad - ICGT (1992) Leaf descriptors. Unpublished data supplied on computer diskette by International Cocoa Genebank, Trinidad, University of the West Indies, Trinidad.